Missouri Botanical Garden A Journal for Botanical Nomenclature VOLUME 21 NUMBER 1 2011 Volume 21, Number 1 April 201 1 Novon, A Journal for Botanical Nomenclature from the Missouri Botanical Garden The mission of the Missouri Botanical Garden is to discover and share knowledge about plants and their environment, in order to preserve and enrich life, Novon publishes short articles whose primary purpose is the establishment of nomencla¬ ture in vascular plants and bryophytes. All articles are peer-reviewed by qualified, indepen¬ dent reviewers. Manuscripts must fully state and justify the reasons for proposing nova. These may include detailed comparisons with similar taxa, short keys to similar taxa, illustrations to similar taxa, and mechanical nomenclature reasons, among others. Manuscripts whose primary purpose is other than establishment of new nomenclature, which usually are longer manuscripts, cannot be accepted for review. These include reviews, revisions, monographs, or other papers that incidentally include nova. Manuscripts must follow the guidelines in the Checklist for Authors. The Checklist may be downloaded from the Garden’s web site, www.mbgpress.info, or authors may contact the man¬ aging editor to request a copy. Novon will not knowingly accept manuscripts that have been simultaneously submitted to other journals for consideration or previously published in some form elsewhere. Victoria C. Hollowed Editor Beth Parada Managing Editor Allison M. Brock Associate Editor Tammy Charron Associate Editor Cirri R. Moran Press Coordinator Ihsan A. Al-Shehbaz Consulting Editor Carmen Ulloa Ulloa Consulting Editor George Yatskievych Consulting Editor Kanchi N. Gandhi Nomenclature Consultant Nicholas J. Turland Nomenclature Consultant Roy E. Gereau Latin Editor Ncrvon is included in the subscription price of the Annals of the Missouri Botanical Garden. Price for 2011: $180 per year U.S.A.; $190 Canada and Mexico; $215 all other countries. Four issues per volume. Postal address: Subscriptions: Annals of the Missouri Botanical Garden, P.O. Box 1897, Law¬ rence, KS 66044-8897; Manuscript submissions: Novon, Missouri Botanical Garden, P.O. Box 299, St. Louis, MO 63166-0299. Web site: http://www.mbgpress.info © Missouri Botanical Garden 2011 © Novon is printed on paper that meets the re¬ quirements of ANSI/NISO Z39.48-1992. Novon (ISSN 1055-3177) is published quarterly by the Missouri Botanical Garden, 2345 Tower Grove Avenue, St. Louis, MO 63110. Periodicals postage paid at St. Louis, MO and additional mail¬ ing offices (USPS #006-777). POSTMASTER: Please send address changes to Annals of the Missouri Botanical Garden, P.O. Box 1897, Lawrence, KS 66044-8897. Information on the contents of Novon is present¬ ed in SciSearch®, Research Alert®, Current Con- tents®/Agriculture, Biology, and Environmental Sciences, APT Online, the CAB Abstract/Global Health database, and the ISI® database. The full-text of Novon is available online through BioOne™ (http://www.bioone.org). Volume 21 NO VON Number 1 2011 Arisaema wangmoense (Araceae), a New Species from China Ming-Tai An Forestry College of Guizhou University, Guiyang 550025, People’s Republic of China. gdanmingtai@126.com Hua-Hai Zhang Guizhou Provincial Wild Animal and Plant Administrative Station, Guiyang 550001, People’s Republic of China State Key Laboratory of Systematic and Evolutionary Rotany, Institute of Rotany, Chinese Academy of Sciences, Reijing 100093, People’s Republic of China. Author for correspondence: linqi@ibcas.ac.cn Tang-Ling Wei Wangmo Forestry Bureau, Wangmo, Guizhou 552300, People’s Republic of China Abstract. A new species of Arisaema Mart. (Ara- between A. wangmoense M. T. An, H. H. Zhang & Q. leaflets, serrulate or denticulate margins, and II (A Red List. The genus Arisaema Mart. (Araceae) consists of reported to occur in China (Li et al.. 2010). western China, in April and May 2009, a previously isaema Nakai (Nakai. 1950) was collected on distinctive when compared with related taxa. For CDBI, GZAC, GZTM, HGAS, HIB, HNNU, IBK, IBSC, KUN, LBG, NAS, PE, SYS, and SZ. The new species is described and illustrated herein. In University during April and May 2009. Lin, sp. nov. TYPE: China. Guizhou: Wangmo Co., Lewang, Shuijingwan, 1100 m, 8 May 2009 (d), M. T. An & T. L. Wei 2009-999 (holotype, GZAC). Figure 1. doi: 10.3417/2009125 Novon 21: 1-3. Published on 7 April 2011. Novon An et al. it an altitude of 1100 m. WCW Red List categon. te limb, the unisexual spadix, the and the anthers that dehisce by Taxonomic Notes on the Genista ephedroides Group (Fabaceae) from the Mediterranean Area Gianluigi Bacchetta Centro Conservazione Biodiversita (CCB), Dipartimento di Scienze Botaniche, Viale S. Ignazio da Laconi 13, 1-09123 Cagliari, Italy, bacchet@unica.it Salvatore Brullo Dipartimento di Botanica, Via A. Longo 19, 1-95125 Catania, Italy, salvo.brullo@gmail.com Tiziana Cusma Velari, Laura Feoli Chiapella, and Vera Kosovel Dipartimento di Scienze della Vita, Via Giorgeri 10, 1-34127 Trieste, Italy, chiapell@univ.trieste.it Abstract. A revision of the taxa belonging to the study, carried out on the basis of the literature, investigations, allows the recognition of 13 taxa. Eight Brullo & De Marco, and five are new to science. Genista bocchierii Bacch., Brullo & Feoli Chiapella, G. subsp. fodinae Bacch., Brullo & Feoli Chiapella, and G. ovina Bacch.. Brullo & Feoli Chiapella are yielding higher counts. In addition to 2 n = 48, the new droides group, IUCN Red Fist, Mediterranean. Novon 21: 4-19. Published on 7 April 2011. presented. According to Valsecchi (1986a, 1986b, 1993a, 1993b), De Marco et al. (1987), Brullo et al. (1993), Brullo and De Marco (1996), Estabrook (2001), De Castro et al. (2002). and Pardo et al. De Marco. On the basis of the literature, as well as Mediterranean species of Genista. According to Spach (1844) and Valsecchi (1993b), the G. herbaria (BC, BM, C, CAG, CAT, FI, G, EG, M, NAP, OXF, P, PAF, RO, TO, TSB, W, WU). These investigations revealed other taxa that are distributed in the central Mediterranean area. occur in the Aeolian and Pontine Islands, respec- doi: 10.3417/2009026 flniiiil et al. {■Iff? ft et al. Novon Volume 21, Number 1 2011 Bacchetta et al. The Genista ephedroides Group (Fabaceae) et al. Novon et al. The et al. et al. et al. Transfer of Anoectochilus tetrapterus to Odontochilus (Orchidaceae) 2011. Volume 21, Number 1 2011 42191) of i A New Species and Combination in Central American Solanaceae \ University of Utah, Salt Lake City, Utah 84112, 1 2011. Novon 25 y forked, with 15 to 30+ fl the tube 1-1.5 mm, the lobes 5-8 X 2-3 mm. mm, the free part 0.5-1 mm; anthers 3-4 X ca. 1.5 parts of Chiriquf Province, fruiting specimens have been collected in February, IIJCN Red List category. According to the IUCN Red List criteria (IUCN, 2001), Solarium sotobos- quense is classified as VU Bla+2a; D1 (Vulnerable; extent of occurrence less than 20,000 km2 and area of 26 27 Typifications and New Synonymies in Capanemia (Orchidaceae, Oncidiinae) Cristiano Roberto Buzatto Universidade Federal do Rio Grande do Sul, Institute de Biociencias, Departamento de Botanica, Programa de Pos-graduatjao em Botanica, Av. Bento Gonyalves 9500, 91501-970, Porto Alegre, Rio Grande do Sul, Brazil, crbuzatto@gmail.com Rodrigo B. Singer Universidade Federal do Rio Grande do Sul, Institute de Biociencias, Departamento de Botanica, Av. Bento Gomjalves 9500, 91501-970, Porto Alegre, Rio Grande do Sul, Brazil. rbsingerl@yahoo.com Gustavo A. Romero- Gonzalez Orchid Herbarium of Oakes Ames, Harvard University Herbaria, 22 Divinity Avenue, Cambridge, Ma-— aclm-ell- 02138, U.S.A. romero@oeb.harvard.edu Cassia van den Berg Universidade Estadual de Feira de Santana, Departamento de Ciencias Biologicas, Av. Transnordestina s/n, 44053-015, Feira de Santana, Bahia, Brazil, vcassio@gmx.de Abstract. Capanemia Barb. Rodr. (Orchidaceae, C. brachycion (Griseb.) Schltr., C. carinata Barb. Rodr., C. gehrtii Hoehne, C. micromera Barb. Rodr., C. supetflua (Rchb. f.) Garay, and C. therezae Barb. Brade. is recognized as valid but not treated here. riograndensis Pabst, and C. spathuliglossa Pabst are referred to C. micromera ; and C. fluminensis Pabst is silla, C. spathuliglossa. and C. therezae. A neotype is total de sete especies. seis das quais sao tratadas Rodr., C. supetflua (Rchb. f.) Garay e C. therezae Brade, e reconhecida como valida mas nao e tratada Pabst e C. spathuliglossa Pabst com C. micromera-. e Schltr. his patron and benefactor (Sa, 2001). Barbosa Novon 21: 28-33. Published on 7 April 2011. doi: 10.3417/2009058 Rodrigues (1877: 137-138) originally described two species (C. uliginosa Barb. Rodr. and C. micromera cm high) with sympodial growth. The LI are long. (Rchb. f.) Garay). The leaves can 1 o 20). The flowers are s, Ausentes, C. ff. Buzatto 486 (ICN); Sao Leopoldo, J. Dutra 1002 (HB, ICN); Tenente Portela, J. L. Waechter 1663 (ICN); Tones. J. L. Waechter 1676 (ICN); Vacaria, C. R. & R. M. Klein 13181 (HBR); Campos Novos, R. Reitz & R. M. Klein 16160 (B. HBR, MBM, NY, PACA, RB); Canoinhas, R. M. Klein 3001 (HB); Capao Alto, C. R. Ponte Alta, R. M. Klein, Le grand. & Marched 9670 HBR); Porto Uniao, ff. M. Klein 3061 (HBR, ICN, K, J aira: Villa Rica, C. A. M^Lindman A1753^( S)_ (MBM); Teresina, P. Dusen 1911 (S); Uniao da Vitoria, H. Rodrigues Jr 1 ingely S G. Hatschbach ill, (MBM); Ventania, ./. Cordeiro 752 (MBM). Rio Grande do Sul: Bento Gonyahes, 4. Sehnem 7211 (PACA); Canela, J. S'. ^nllTc 4Sl"lCNf !FCs18a Castelhano, C. ff. Buzatto 29 (ICN, RSPF); Montenegro, A. Sehnen 4019 (B); Passo Fundo, C. ff. Buzatto 525 (ICN); Porto Alegre, C. Orth 1932 (B); Putinga, T. Klein 2002 (Herbario do Museu de Ciencias Naturais da UNIVATES); Rio Pardo, J. Dutra 41 (ICN); Santa Cruz do Sul, J. L. Kaechte, 1789 (HAS, II N), Sao Francisco de Paula, G. Grazztotm et al, 3547 (MBM); Sao Leopoldo, J. Dutra 1119 iberrtz,\ ?;^ff":: as set NY, S); Bom Retiro, ff. Reitz 6038 (HBR); Campos Novos, ff. M. Klein 4235 (HBR, MBM, RB); Catanduvas, L. B. 4&80*'mn7 Yb86 (BffHM ' aY)''S^mn 53^^^ 1168 .anemia dmeniana (Kraenzl.) Porto & Brade, Rodrigue- sia 1: 20. 1935. Basionvm: Quekettia dmeniana Kraenzl., Ark. Bot. 16(8): 24. 1921. TYPE: Brazil. Parana: Curitiba, s.d., P. Dusen & ff. Lange 8529 1,8 41003, K as Capanemia therezae. following Ret 60C.l(a) in the International Code Taxonomic Notes for Salvia aucheri (Lamiaceae) from Southern Anatolia, Turkey Celep et al. Leaf dimensions (cm) (1.4-)4-13.5 l-3.5(-4.5) ators of ANK, BM, E, G, GAZI, HUB, ISTE, and K herbaria for allowing us to study their Salvia collec- n & F. Celep 999 (ANK); A A New Species of Hagenbachia (Anthericaceae) from Ecuador City, Iowa 52240, U.S.A. Volume 21, Number 1 2011 Novitates Agrostologicae, II. Parodiolyra aratitiyopensis, un Nuevo Bambu Herbaceo del Sur de la Guayana V enezolana Jose Ramon Grande Allende Volume 21, Number 1 2011 Aegiphila brenesii , a New Name for the Long-lost (Right Before Our Eyes) 2011. oo-o-ola, Validation of the Genus Name Darcyanthus and Its Sole Species, Darcyanthus spruceanus (Solanaceae) doi: 10.341: 2011. Three New Costa Rican Species of Calathea (Marantaceae) from Montane Wet Forests Volume 21, Number 1 2011 Kennedy Calathea (Marantaceae) from Costa Rica mu to peach-colored basally with apical petaloid portion purplish or dark purplish pink. ca. 22 X 7 mm; (LC) IUCN Red List criteria (IUCN, relatively large (3.2-4 X 4-5.5 cm), p 6 Sep. 1972, 1L Kennedy 1558 (BM. CR, DUKE,' INB, MCE US); Bajo La Hondura, 10°04'N, 83°59'W, 8 Sep. 1976, H. H. Kennedy 4585 UR. INB, MO), Paique Nac Biaulio Brauho Carillo, La Montura, along ridge, 25-30 July 1982, C. Todda 1958 (CR). it herb, 0.9-1. 9 m high, b confluent. (2.1-)3-8(-9.5) cm; petiole dark green or Volume 21, Number 1 2011 Novon basally grading to yellow near apex, appressed long in subtending cauline leaf, (39-)104-186(— 236) cauline leaf 20-38 cm. others (27-)62-113 Inflorescences 1 to 5 per shoot, the first terminal, subtending leaf, strongly complanate, rectang 9.2-13 X (3.2-)5.5-8 cm. width substantially let side. 44-97(— 111) cm; bracts 8 to 18. distichous, refuse, often dying back at apex, 3. 8-4.5 X 2.8 3.5 pairs, outer surface of bracts deep olive-green, except prophyll firm centrally, thickened along the base of apical 1/5 of center back and carina tomentose, glabrous basally, 3. 5-3.9 X 1.2-1. 3 cm, 0.5-0. 8 cm linear, apex acute, white basally, very tip tinged acute to 90°. white basally^ apex tinged faint but always with tuft of hairs at very apex (X14 magnification), hairs to 0.5 nun, 17-20 X 2-4 mm; X 4-5 mm; outer staminode broadly elliptic, clawed at base, rounded, white, faint yellow to yellow-green, with one side wider than the other, apex off-center, 11-12.5 X 7-8 mm, crowned by a persistent calyx; seeds usually 3 per capsule, trigonous, blue, ca. 8.5 X 3.5 X 3.5 mm, bearing a white aril. sis occurs at mid-elevations, front 845 to ca. 1500 nt, thea tarrazuensis should be considered as Data Deficient (DD) according to IUCN Red List criteria (IUCN, 2001), because it is only known from a same time lessens the threat. Etymology. The specific epithet, tarrazuensis, lengtluwidth ratios of 2.13-4.8:1 and an "acuminate- peduncles (44— 97[— 111] cm), the bracts longer than 57 o store and study all the M > and use of the 1 comments. C. Niezgoda provided accommodations and ^transport for my stay at F and T. Salvato provided Plantas de Costa R BRIT. CAS. DAV. DH. DUKE. F. INB. MO. NY. PMA. SCZ. SEL. UC. US. and WIS for loan of their specimens and the curators of AAU. CR. GH. MO. UCR. and USJ for the use of their facilities. Thanks Calathea matudae and C. sousandradeana , New Species of Marantaceae from Mexico and Northern Central America Novon Volume 21, Number 1 2011 facilities: G. Davklse (MO), C. Niezgoda (F), G. Salizar (MEXU), E. Poll (UVAL), M. Veliz and J. species. I thank the curators of BRIT, CAS, DAV, DH, DUKE, F, MEXU, MICH, MO, NY, TEX, WIS, UC, and US for loan of their specimens. Calathea leonoriae (Marantaceae), una Nueva Especie de Veracruz, Mexico o Postal 63, Xalapa 91070, Vei o Postal 63, Xalapa 91070, Vei Volume 21, Number 1 2011 et al. Validation of the Name Asparagus kansuensis (Asparagaceae) Qi Lin and Zhi-Rong Yang State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, People's Republic of China. Author for correspondence: zry@ibcas.ac.cn Wang & T. Tang ex S. C. Chen (Asparagaceae) from China was invalidly published in 1978, because two 1978 diagnosis. Thus, the original ascription to K. S. Hao 416 (Chen, 1978). We examined sheets at PE, all attributed to the type collection K. S. two gatherings from different plants are represented by Hew 416 and these separate specimens were simultaneously indicated as type (Chen, 1978). Thus, Articles 37.1, 37.2, 8.1, and 8.2 of the International Code of Botanical Nomenclature (McNeill et al., 2006). The mixed gathering cannot be selected as lectotype. viz. Articles 9.2 and 9.15, or regarded as syntypic (Art. 9.4). Without consideration of the validity of the name, Chen and Tamanian (2000) accepted Asparagus original publication and by indicating one of the authorship of this name remains F. T. Wang & T. ascribed to them, according to Article 46.2 of the Neill et al., 2006). Asparagus kaiisuensis F. T. Wang & T. Tang ex S. Wang & T. Tang ex S C. Chen, Acta Phytotax. Sin. 16(1): 94, nom. inval. TYPE: China. Gansu: Wenxian, 910 m, 15 June 1930 (c? fl.), K. S. Hao 416a (holotype, PE 00034519). Chen (1978: 94). ^ Para-types. CHINA. Gansu: Wenxian, 910 m, 15 June 1930 ( 9), K. S. Hao 416b (PE 00034519), ( 9), K. S. Hao 416c (PE 00218312), ( 9 ), K. S. Hao 416d (PE 00218311); Wenxian, (9), /. Y. Zhang 7085 (PE). Virtual Herbarium Establishment (grant number 2005DKA21401). Sinica. Acta Phytotax. Sin. 16(1): 91-96. L. Pp. 208-215 in Z. Y. Wu & P. H. Raven (editors). McNeill, J., F. R. Barrie, H. M. Burdet, V. Demoulin, D. L. Hawksworth, K. Marhold, D. H. Nicolson, J. Prado, P. C. Silva, J. E. Skog, J. H. Wiersema & N. J. Turland doi: 10.3417/2009077 Novon 21: 69-70. Published on 7 April 2011. Novon Validation of Two Names, Berchemia omeiensis and Rhamnus kwangsiensis (Rhamnaceae) Qi Lin and Zhi-Rong Yang State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, People's Republic of China. Author for correspondence: zry@ibcas.ac.cn omeiensis W. P. Fang ex Y. L. Chen & P. K. Chou and Rhamnus kwangsiensis Y. L. Chen & P. K. Chou (Rhamnaceae), were published invalidly in 1979 neously indicated. Both plant names are validly mia omeiensis W. P. Fang ex Y. L. Chen & P. K. Chou (Chen & Chou, 1979a) and Rhamnus kwang¬ siensis Y. L. Chen & P. K. Chou (Chen & Chou, 1979b) (Rhamnaceae), were not validly published clciture (McNeill et al., 2006), the two names were publication, Chen and Schirarend (2007: 129, 149) Flora of China. The current paper validates the two original publications (Chen & Chou, 1979a, 1979b) Y. L. Chen & P. K. Chou because the name and the ascribed to them (cf. Art. 46.2, McNeill et al., 2006). B< I ensis W. P. Fang ex Y. L. Chen & P. K. Chou, sp. nov. Berchemia omeiensis W. P. Fang ex Y. L. Chen & P. K. Chou, Bull. Bot. Lab. N. E. Forest. Inst., Harbin 5: 16. 1979, validating Latin diagnosis appeared in Chen and Chou (1979a). Paratypes. CHINA. Chongqing: Chengkou, T. L. Dai 100149 (PE [2]), 100280 (PE [2]), 100666 (PE); Fengjie, C. Y. Chang 25036 (PE), 25459 (PE), 25999 (PE); Nanehuan, G. F. Li 60889 (PE), J. H. Xiong & Z. L. Zhou 92428 (PE). Guizhou: Tongzi, Y. Tsiang 5141 (PE [2]). Hubei: Jianshi, W. B. Lin 92 (PE); Xuanen, II. J. Li 2760 (PE), 3271 (PE). Sichuan: Mt. Emei, Y. L. Chen 33 (PE [2]), H. L. Tsiang & S. S. Chang 30619 (PE), 31674 (PE), G. H. Yang 5456.3 (PE), 55157 (PE), 5600.3 (PE), 5725.3 (PE). Rhamnui k\\ uii"sieiisis 1 L. Chen & P. K. Chou, K. Chou, Bull. Bot. Lai)' N. E. Forest Inst., Harbin 5: 77. 1979, nom. inval. TYPE: China. 11111 ( holotype, IBK). Discussion. X. F. Deng 11111 (IBK) is chosen as with the protologue, and was annotated as “TYPUS” validating Latin diagnosis appeared in Chen and Chou (1979b). Paratypes. CHINA. Guangxi: Guilin, Yanshan, ,Y. F. Deng 10948 (IBK), Exped. Guangxi 4100 (PE [2]), C. H. Tsoong 808475 (IBK, PE); Linggui, Exped. Sine- German 341 (IBK); Yongfu, S. Y. Li 6021 (IBK); Liangfeng, Z. N. Deng 13578 (IBK); Dabu, Z. N. Deng 13642 (IBK); Yangslmo, Z. Z. Chen 5.3210 (IBK). by the Plant Specimen Digitization and Chinese 2005DKA21401). nom. inval. TYPE: China. Sichuan: Mt. Emei. 25 May 1957, G. H. Yang 54723 (holotype, PE 00023541). Discussion. G. H. Yang 54723 (PE) is chosen as with the protologue, and was annotated as “TYPUS” Y. L. & P. K. i Bull. Bot. Lab. N. E. Forest. Inst., Harbin 5 Chen, Y. L. & P. K. Chou. 1979b. Materiae ad iloram Rhamnaceamm sinicamm. Bull. Bot. Lab. N. E. Forest. In 5; 73-94. doi: 10.3417/2009015 Novon 21: 71-72. Published on 7 April 2011. Novon Berchemia Neck, ex" DC., Rhamnus L. Pp. 124-130, 139-162 in Z. Y. Wu & P. H. Raven (editors). Flora of McNeill, J„ F. R. Banie, H. M. Burdet, V. Demoulin, D. L. Hawksworth, K. Marhold, D. H. Nicolson, J. Prado, P. C. Silva, J. E. Skog, J. H. Wiersema & N. J. Turland Two New Species of Machaerium (Leguminosae) from Bahia and Southeastern Brazil Carlos Victor Mendonga Filho Programa de Pos-Graduayao em Biologia Vegetal, Institute) de Biologia, Universidade Estadual de Campinas, CP 6109, 13083-970, Campinas, Sao Paulo, Brazil. Current address: Departamento de Ciencias Biologicas, Universidade Federal dos Vales do Jequitinhonha e Mucuri, Campus JK, Rodovia MGT 367, Km 583, n° 5000, Bairro Alto da Jacuba, 39100-000 Diamantina, Minas Gerais, Brazil, cvmendonca@gmail.com Haroldo Cavalcanti de Lima Institute de Pesquisas Jardim Botanico do Rio de Janeiro, Rua Pacheco Leao 915, 22460-0.30, Rio de Janeiro, Brazil, hlima@jbrj.gov.br Eli.ana R. Forni-Martins and Ana M. G. A. Tozzi Departamento de Biologia Vegetal, Institute de Biologia, Universidade Estadual de Campinas, CP 6109, 1.308.3-970, Campinas, Sao Paulo, Brazil, elianafm@unicamp.br; anatozzi@unicamp.br rium Pers. sect. Oblonga (Benth.) Taub. (Legumino- macaense C. V. Mendonga, A. M. G. Azevedo & H. Ecologica Macae de Cima, an area of Atlantic forest differs by its smooth bark, smaller leaflets, sunken Rudd, from which it differs by bark characters, the bracteoles that are shorter than the calyx, and its glabrous leaflets, ovate bracteoles. and glabrous Resumo. Duas novas especies brasileiras de Ma- inacaense C. V. Mendonga, A. M. G. Azevedo & H. de Macae de Cima, uma area de Floresta Atlantica no imersas, bracteolas estreitas, e estandarte menor. Ela e tambem similar a M. hatschbachii Rudd, da qual Minas Gerais, Espfrito Santo e Bahia, ocorrendo nas Florestas Atlantica e mesofftica, ou na vegetagao Benth.) Ducke da qual se difere pelos folfolos thum Spruce ex Benth. uma liana ou arbusto da Amazonia, com inflorescencias paniculadas. rnais 130 species with all but one species (M. lunatum (L. doi: 10.3417/2009029 Novon 21: 73-77. Published on 7 April 2011. Novon Volume 21, Number 1 2011 Mendonga Filho et al. Machaerium (Leguminosae) from Brazil oblong, 0.5-1 .8 X 0.2-0. 6 cm, petiolulate or sessile, axillary or terminal panicle. 4-11.5 cm; bracts lanceolate, ca. 3X1 mm; calyx 3-4 X 2-2.5 mm, fulvous-sericeous; standard ca. 6 X 4 mm, adaxially white-sericeous; petals lilac, keel petals 6-6.5 X 1.5- monadelphous to diadelphous (the vexillary stamen style straight, 2.5-3 mm; ovary ca. 2.5 X 0.5 mm. Fruit a 1-seeded samara, 3.8-6 X 1.2-1. 3 cm; seed chamber 1.3-1. 5 X 0.8-1 cm. State; it grows in the upland Atlantic forest of the 1200 m. IUCN Red List category. Machaerium macaense according to IUCN Red List criteria (IUCN, 2001). Ecologica de Macae de Cirna. tion of Taubert (1891). Machaerium macaense assigns similar to M. nyctitans (Veil.) Benth., differing from this taxon by its smooth bark, smaller leaflets (vs. narrower bracteoles (vs. 3-5 X 0.5-1. 5 mm), and smaller standard petal (vs. 6.4-10 X 5-8 mm). The which it differs in bark characters, the bracteoles that Paratypes. BRAZIL. Rio de Janeiro: Nova Fnburgo, 3830 (F, P, RB, U), H. C. de Lima, F. C. Garcia & G. Laurentino 5024 (MO, RB), C. M. B. Correia, S. V. A. Pessoa , J. Caruso, L. C. Siha & J. C. Siha 99 (CEPEC, NY, RB), A. M. S. F. Vaz, L. Syhestre, C. M. Vieira & L. C. Silva 718 (NY, RB), C. V Mendonga & E. S. Camara 616 (UEC). M. G. Azevedo,' sp. nov. TYPE: Brazil. Espfrito 2 of rd. Bicmba, 19 Sep. 2001, D. A. Foil 4060 (holotype, CVRD; isotype, MO). Figure 2. Shrubs or small trees to 3 m tall, 5-10 cm DBH. striate, spinose, 4-4.5 X 1.3-1. 5 mm; rachis 0 6-1.3 X 0.2-0. 4 cm. petiolulate or sessile, to tomentose: petals cream, standard 5-6 X 4-4.5 keel petals 5-5.5 X ca. 2 mm; wing petals 5-6 X 1.5- 2 mm: stamens 10. monadelphous. 3-4.5 mm, the free part of filaments 1.5-2 mm; style straight, 1.5-2 4-4.5 X ca. 1 cm; seed chamber ca. 1 .2 X 0.8 cm. num is found in southeastern and northeastern Brazil, grows in lowland Atlantic and mesophitic forests, or IUCN Red List category. Machaerium jobimia- insufficient information is available to apply IUCN Red List criteria (IUCN, 2001). September. 1994). multifoliolate leaves, oblong leaflets, and spinescent Novon Five New Species of South American Gentianella (Gentianaceae) u Ontario L8N 3H8, Canada, jpringle@rbg.ca 2011. doi: 10.341" Volume 21, Number 1 2011 Novon (22-)30-40 mm diam.; corolla lobes elliptic-rhombic to rhombic-obovate, (10-)12-18 X (5-)10-13 mm, 3.5-4.5X as long as tube, 0.9-1. 5X as long as wide, of length: anthers purple. 1.4-2. 5 mm: ovary with ca. 1.8 X 1.4 mm. Capsule not seen. tianella decemnectaria were collected at 3120-3500 Discussion. Hagen and Kadereit (2001, 2002) Comastoma (Wettst.) Toyok. and Lomatogonium A. Catharanthus roseus (L.) G. Don rather than being are more or less rhombic; that is, the margins, rather only slightly convex distally. Although the filaments out Gcntianella . the relatively long filaments of G. are basally connate-sheathing. Distally the sheaths inflorescences they are absent, not even a transverse In its decumbent stems, the size and shape of its is similar to G. kuntzei (Gilg) T. N. Ho & S. W. Liu, of are 5-15 mm, shorter and more slender than those of is only 12-18 mm when closed, with narrowly floral trichomes of G. euphorbiifolia. in which a summit of a lateral flange on each side of the 9-15 mm, tube 45-6.5 mm, ridged along commis- r, 45-9.5 X 2.3-4 mm (width a late, 15-25 mm long, probably opening to ca. 1 cm diam.; corolla lobes obovate, 10-16 X 5-9 mm, 2.5- ■s 5, ca. 1.5 X 1.2 mm; 2-2.5 X 2-2.5 n ii (Gilg) Fabris ex T. N. Ho & S. Novon G. 18 Oct. 1984, 1 G. Beck 8936 (HAM, Novon Gentianella. the combination of basal leaves in a lobing, and color to those of G. brunneotincta (Gilg) J. S. Pringle, of northern and central Pent, and G. hirculus (Griseb.) Fabris, of Ecuador. Plants of those 4 mm long. In G. e lobes proportionately narrower than those of G. Novon unless^ near the base, and the ealyx lobes are shorter Chuco, Pampa de la Julia (al este de Quimvilea), 23 May 2001, S. Leiva G. & P. Leiva 2574 (F not seen, HAM, HAO 7°55.3'S,' 78°10.2'0, 23 May 2001,’ A. Sagmtegui A. & M. E. Zapata C. 16576 (F not seen, HAM, HAO not seen). with 1 pair of Novon Himalayopteris , a New Fern Genus from India and the Adjacent Himalayas (Polypodiaceae, Polypodioideae) Wen Shao and Shu-Gang Lu Institute of Ecology and Geobotany, Yunnan University, Kunming, Yunnan 650091, People's Republic of China, shaowenl9792005@163.com Abstract. A new fern genus. Himalayopteris W. dia, IUCN Red List. Nepal. Polypodiaceae. later treated as Goniophlebium erythrocarpum (Mett. ex Kuhn) Bedd. by Beddome in 1876. When description of Beddome (1876, 1883) in having a Presl to the genus Phymatodes C. Presl (Ching, 1933), and then to Phymatopsis J. Sm. (Ching, 1964), in what is now the subfamily Selligueae in the differs from that seen in Goniophlebium. From HITBC, we confirm that the characteristics of the as shown in Figure ID. Nonclathrate rhizome scales Phymatodes and Phymatopsis, Pichi Sennolli (1973) ris erythrocarpa (Mett. ex Kuhn) Pic. Semi., but Japanese botanists treated Phymatopteris Pic. Semi, as a synonym of Crypsinus C. Presl (Nakaike, 1987). Rodl-Linder (1990) accepted this synonymy, with Goniophlebium. Lu (2000) supported the prior treatment of Pichi Sennolli (1973), while Hovenkamp (1998) further treated Phymatopteris and Crypsinus as synonyms of Selliguea. Fraser-Jenkins (2008) sepa¬ rated this and certain other species from Selliguea. as the genus Piehisennollia Fraser- Jenk. (non Piehiser- Jenk. (Fraser-Jenkins, 2009). However, until now no one has separately recognized the individual species Mehra (1961: 162) as x = 37 (cited by Liive et al., 1977: 57, as 2n = 74). This basic number is prevalent Ching). The goniophlebioid venation pattern and from Phymatopteris and prevent the inclusion of this this distinctive taxon at the genus level. Novon 21: 90-93. Published on 7 April 2011. doi: 10.3417/2009115 f I I 1 i i K). Figure 1. i of its sole known species, Hima- & Lu Fraser-Jenkins, C. R. 2009 [2010], A brief comparison of ‘.Scril .XiSi Rubiacearum Americanarum Magna Hama Pars XXVI: New Species of Hoffmannia (Hamelieae) and More Comments on the Genus Charlotte M. Taylor and Roy E. Gereau Mi— ouri Pol ;im leal On iilc’ii, P.0. Box 299, St. Loui-, \li-omi (.3 K.6-0299, U.S.A. charlotte.taylor@mobot.org; roy.gereau@mobot.org Abstract. Species of Hoffmannia Sw. (Rubiaceae, montane areas of Central America and southern five newly described species and all the Hoffmannia their leaves are: //. formicaria C. M. Taylor of Costa corollas with the acute to acuminate lobes longer than western Panama, which differs from //. v esieulifera that are rounded at the base and its epiphytic habit; from H. obovata (Ruiz & Pav.) Standi., H. latifolia H. coriacea C. M. Taylor of central Peru is the lobes; //. costari-censis C. M. Taylor of Costa Rica are acute in bud; //. pacifiea C. M. Taylor of Costa //. nieotianifolia, (M. Martens & Galeotti) L. 0. Williams in its larger red to purple flowers and larger the base and have numerous secondary veins plus its Resumen. Las especies de Hoffmannia Sw. (Rubia¬ ceae, Hamelieae) se concentran en areas premonta- Nicaragua. Costa Rica. Panama y el noroeste de Novon 21: 94-117. Published on 7 April 2011. doi: 10.3417/2009118 96 Novon this region, and Deppea Cham. & Schltdl. (Lorence & Dwyer. 1988). Hoffmannia (Burger. 1999), and Hamelia Jacq. (Elias, 1976) have species-level (Benth. & Hook, f.) Bremek. (Psychotrieae), which is pleura can be separated by its pseudoaxillary node vs. consistently in both axils in Hoffmannia ), 5- Morphology of Hoffmannia Plants of Hoffmannia are often quite succulent and likely that the same is true for 'Hoffmannia , but field of a single plant. The full range of this variation has not been documented in many descriptions. mannia species (Dwyer, 1969; Burger, 1999) and has Rubiaceae (Dwyer, 1969; Burger, 1999), in several Hoffmannia species with apparently broad ranges of including field observations. No flower odor has been The comparative length of the corolla tube versus species of Hoffmannia , as noted by several authors (e.g., Standley, 1934). Hoffmannia flowers with the corolla lobes much longer than the tube and widely (1934, 1936, 1938), Dwyer (1969), am:f Burger same plant. The basis for Delprete’s (1998) unique atic for his generic comparison there. Dwyer (1969) noted variation in stamen number in Volume 21, Number 1 2011 Taylor & Gereau New Species of Hoffmannia merits study. Dwyer (1969) and Hayden (1968) also noted 4-locular ovaries (//. cuneatissima B. L. Rob., //. Hoffmannia' s fruits are generally fleshy, though of juicy. The fruits are usually baccate and dispersed noted that sometimes these fruits are tardily release the numerous tiny dry seeds. Dwyer (1969) not be the mature fruit color. However, some (1968), but the seeds of most Hoffmannia species Taxonomic Concepts and Biogeography At least two thirds of the species of Hoffm annia are Rubiaceae genera (Williams, 1973; Burger, 1999). contrast, the species of Hoffmannia found in South mark, 1974). Two species are found on Isla del Coco in the Pacific Ocean (Trusty et al., 2006), and both Kiehn (1995) noted that polyploidy is found in the Hoffmannia (Kiehn, 1986). He also documented The Hoffmannia vesiculifesa Group corollas. (The specific epithet of //. vesiculifera spelling “vesciculifera.” but that spelling has not the reference “Words by William Whittaker” pandurate leaf blades. These leaf blades are narrowed then truncate at the base of the blade tissue with this Volume 21, Number 1 2011 Taylor & Gereau New Species of Hoffmannia methodology. The ic and floristic: the and no field studies have been done targeting the on incomplete survey of the available data, which are essential to understanding the threats to i criteria (IUCN, 2001) based on the totality of our in the form of a map and the calculated f “Show Detail”). The assessment parameters were calculated using the IUCN Rating tool (Moat, 2007) in ArcView CIS 3.2 (ESRI, 1999), with the grid cell size used for calculating Area of Occupancy (A00) These assessments are not being submitted to IUCN for publication on the Red List (). and the basis for these should be carefully evaluated by the Cuatrecasas (1953) and Steyermark (1966) distin- two stipulelike bracts, and stipules partially connate at the base. However, detailed study of the group revealed that the three latter characters also occur in C. M. Taylor is here lowered to a variety of A. eduli-s (Rich.) A. Rich, ex DC., as the original differentiating characters are not complete when considering the Dwyer. Ann. Missouri Bot. Card. 67: 46. 1980. TYPE: Panama. Colon: 2-3 mi. up Rio Guanche, 10-20 m, 19 Jan. 1973 (fr.). H. Kennedy & R. Foster 2197 (lectotype. designat¬ ed by Lorence [1999: 19]. MO 2163702 [sheet 2 of 2]; isotypes. F. MO 2163702 [sheet 1 of 2]. PMA not seen). MO 2163702. and apparently Dwyer (1980) had ♦This s 136 Novon distributed from Nicaragua to Panama, and charac¬ terized by leaf blades with (nine to)ll(or 12) pairs of is closely related to A. dwyeri Delprete & C. H. short petioles ([5-J10-18 mm long vs. [5— ]10— 27 seems to occur in lowland forests, below 400 m, 600-900 m altitude. Selected specimens examined. COSTA RICA. Heredia: (MO). NICARAGUA. Rio San Juan: Reserva Indio-Maiz, 10°46'N, 83°53'W, 100-200 m, R^Rueda et al. 8871 (MO). PANAMA. Colon: Rio Guanche, ca. 2 km upstream from the elevation note “2700 ft.” is lacking on the that the holotype was collected by Duke and Dwyer, an error that was corrected by Dwyer (1980) in his treatment of Rubiaceae for the Flora of Panama. The 2005), specialist in Neotropical Rubiaceae (and who and Panama (for his biography, see Croat et al., 2006i. of cloud forests of Panama, at 600-900 m altitude. It elliptic or slightly obovate leaf blades, but differs by abaxially (vs. velutinous or hirsutulous abaxially), the glabrate), and the dentate or jagged calyx margin (vs. Llano Grande, 600 m, Antonio 3605 (MO); Cerro Pilon, Motel, Croat 14779 (F, MO, NY). 2. Alibertia dwyeri Delprete & C. H. Perss., nom. Dwyer, Phytologia 17: 446. 1968. non Alibertia. 1927: 123 '^= Alibertia edulis (Rich.) A. Rich, ex DC., Prod. (DC.): 4: 443. 1830.]. TYPE: Panama. Code: Cerro Pilon [Pilon], El Valle, 4 Jan. 1968 (fr.), J. A. Duke & B. R. Lallathin 15014 (holotype, MO 1968459; isotype, MO 2163702). collected at Cerro Pilon at 2700 ft. [ca. 823 m] by B. R. Lallathin in 1968, coll. no. 15014. According to an additional label added by C. M. Taylor, the original label was probably mistyped, as the digit “1” seems to be added afterward in front of “5014.” Also, 3. Alibertia edulis (Rich.) A. Rich, ex DC., Prod. (DC.) 4: 443. 1830. Basionvm: Genipa edulis Rich., Actes Soc. Hist. Nat. Paris 1: 107. 1792. TYPE: Guyane Francoise [French Guiana]. S. loc„ 1792, Le Blond s.n. (lectotype, designated here, P). type of this taxon was collected by L. C. Richard. Blond in French Guiana and described shortly after Rich., it is impossible that he collected the type of Botanical Nomenclature (McNeill et al., 2006: Art. 9.9). Taylor et al. Taylor et al. Osa, Reserva Forestal de Golfo Dulce, fila entre Rancho Quemado & Drake, bosque alto a la par 2003, B. Hammel, I. Perez & J. Salazar 22841 (holotype, INB; isotypes, CR, USJ). Fi^tre 1A, B. :> elliptic, 10.5-25.5 X 2-5.8 140 Novon terrestrial habit with the stems unbranched, its (Taylor. 2001). This new species is only known from the Osa Peninsula in the Golfo Dulce region, an Costa Rica. These small understory plants are rare. both axils at each node, and blue to black fruits adaxial surface. Paratypes. CObTA RICA. Puntarenas: canton Osa, Kennedy & J. Solomon 17865 (t 'It, F, MO), B. Hammel, M. M. Chavarria. P. Maas & H. Maas 18067 (CR, F, MO), 18287 (CR, F, INB, MO). Bolivia and northern Argentina (Taylor, 1997). better understood (Bremer & Eriksson. 2009). used here generally follow Taylor (1997). Keys to the Fila Matama, Valle de La Estrella. El Progreso, cabeceras del no Cariei, 9°47'N. 83°08'W. 1400 m. 26 Apr. 1989. G. Herrera & A. Chacon 2805 (holotype, CR; isotypes. INB. M0- 05016981). Figure 2A. B. Leaves paired; blades elliptic. 10.5-17 X 4-6 cm. hirsute to hirtellous along veins; secondary veins 9 to II. a Taylor et al. Taylor et al. 146 Novon and its corolla lobes that terminate in a linear smaller (up to 15 X 18 cm) and generally branched similar to P. megistophylla Standi, of western and its corolla lobes that terminate in a linear appendage 0.5-1. 5 mm long (Taylor & Monsalve. dor. Peru, and Brazil. P. acreana K. Krause and P. ceronii C. M. Taylor, but both of these South leaf veins that are broadly looping and only weakly Replaced synonym: Rudgea thrrsiflora Donn. Sm., Bot. Gaz. 61(5): 375. 1916. non Psychotna thrrsiflora Ruiz & Pav., 1799. TYPE: Costa m. Apr. 1895. A. Tondu z 9579 (holotype. US 943477; isotype. BR). ea Salisb. based apparently on its stipules, which thrrsiflora was classified there in Rudgea ser. initially, i.e., on the youngest stipules at the stem apex (Taylor et al.. 2004). These glandular append - interpetiolar portion of the sheath (vs. medially or basally). The stipules of R. thrrsiflora are united around the stem into a truncate sheath and have two bristles borne adaxially (i.e.. on the inside) near the Rudgea thrrsiflora is thus anomalous within Rudgea black at maturity indicate its classification in to Rudgea (Robbrecht & Manen, 2006)! Accordingly. locality. the flowers sessile in small groups, relatively small immature then turn black and have five (rather than inflorescences and its longer cc irollas (with the tube paniculiform inflorescences and shorter corollas (with the tube 2.5-3 mm long) in P. racemosa. Psychotna the western Amazon basin, but P. tsakiana is so far by its longer calyx limb ( 0.8—1 mm long) and its larger corollas (with the tube 2.5-3 mm long), versus the calyx limb ca. 0.5 mm long and the corollas with the tube 1.5-3 mm long PANAMANIAN SPECIES* front eastern Panama. IV. Bristan 1187 (MO). The not explained, and at that time Bathysa had not been reported from Central America (Standlev. 1938) or ♦This st Taylor et al. Arundinella kerrii and Dimeria kerrii , Two New Endemic Species from Thailand (Poaceae, Panicoideae) Atchara Teerawatananon Natural History Museum, National Science Museum, Technopolis, Pathum Thani 12120, Thailand. teerawaa@gmail.com Sarawood Sungkaew Department of Forest Biology, Faculty of Forestry, Kasetsart University, Bangkhen, Bangkok 10900, Thailand; Center for Advanced Studies in Tropical Natural Resources, Kasetsart University, Bangkhen, Bangkok 10900, Thailand. Author for correspondence: fforsws@ku.ac.th Trevor R. Hodkinson School of Natural Sciences, Botany Building, Trinity College Dublin, University of Dublin, Dublin 2, Ireland. hodkinst@tcd.ie and Dimeria kerrii Teerawat. & Sungkaew (Poaceae, Panicoideae) are validly published for the first time. used by Hambananda (1990, unpublished thesis) and Arl wdlxella Raddi TYPE: Thailand. Nakhon Phanom: Tha Uthen, ca. 200 m, 16 Feb. 1924, A. F. G. Kerr 8474 (holotype, BK; isotypes, BM, K). Figure 1. Perennial, loosely tufted. Culms 30-120 cm tall, nous, ciliolate, 0. 5-0.6 mm, with a dense row of hairs 50(— 70) X 0.3-0. 8 cm. glabrous on both surfaces. hairs. Panicles contracted, 5-15 cm long; secondary Spikelets grayish green, ovate-oblong, 3-3.6 X 1-1.5 glumes ovate, 2.5-3 X 0.8-1. 2 mm, acute, 3- to 5- upper glumes ovate to ovate-oblong, 2. 5-3. 5 X 1-1.2 or minutely bifid, shortly awned from the sinus, awns 1.8-2 mm, acute; callus pubescent, hairs 0. 2-0.3 mm; IUCN Red List category. Using the IUCN Red Fist criteria (IUCN, 2001), Arundinella kerrii could be considered as Endangered (EN) or possibly Extinct (EX). The taxon is known only from collections made it. However, insufficient data exist regarding its doi: 10.3417/2009033 Novon 21: 149-153. Published on 7 April 2011. 150 Novon shorter callus hairs that are ca. 1/5 the length of effectively published, by Hambananda (1990: 28-30) in an unpublished thesis (McNeill et al.. 2006: Art. Code of Botanical Nomenclature (McNeill et al.. Volume 21, Number 1 2011 Teerawatananon et al. New Poaceae from Thailand 151 2. 6-3. 5 mm lone n (1/3 — 1/2 length of upper Paratrpe. THAILAND. Nahkon Phanom: Chaiyaburi, ca. 200 m, 1 May 1932, A. F. G. Kerr 21330 (BK, BM, K). Dimeria R. Br. kerrii Nanakom & Norsangsri, 2001, nom. nud. TYPE: Thailand. Satun: Ban Tola Tai [Tola], ca. 50 m, 3 Jan. 1928, A. F. G. Ken- 13868 (holotype, K 648084; isotypes, BM 928281, K 648258). Figure 2. noded at base. Leaf sheaths overlapping below, 6-9 10-20 X 3-4.5 mm. tomentose on both surfaces. Racemes (2 or)3, 8-16 cm, rhachis flattened, 0.6-0. 7 flattened, 0.8-1. 2 mm, margins glabrous, clavate; glumes oblong, 5-5.5 mm, acuminate, keeled, ciliate on keel; upper glumes oblong-elliptic, 5.5-6 mm, margins, broadly winged all along the keel, wings altitude of ca. 50 m. IIJCN Red List, category. Using the IUCN Red List criteria (IUCN, 2001), Dimeria kerrii could be in peninsular Thailand. Because data regarding its the status as Data Deficient (DD). 13868 collection at Kew, but Hubbard never Norsangsri, 2001), but a Latin diagnosis was not accordance with Art. 36.1 and Rec. 36A of the 152 Novon This species is distinguished from all other species (1877-1942), the Irish doctor and h 1 = r f i: 153 staff of the following herbaria: BKF, C, E, K, and NY for the use or loan of specimens. Thanks to Pranom Fund (TRF)/ ^Species sS son, J. Prado, P. C. A New Species of Sciaphila (Triuridaceae) from Hainan Island, China Han Xu Research Institute of Tropical Forestry, Chinese Academy of Forestry, Longdong, Guangzhou 510520, People's Republic of China; Research Institute of Forest Ecology, Environment, and Protection, Chinese Academy of Forestry, Reijing 100091, People's Republic of China. hanxu8 1 @gmail.com Yi-De Li Research Institute of Tropical Forestry, Chinese Academy of Forestry, Longdong, Guangzhou 510520, People's Republic of China Huan-Qiang Chen Jianfengling National Nature Reserve, Jianfengling, Ledong 572542, People's Republic of China Abstract. Sciaphila jianfenglingensis Han Xu. Y. D. Li & H. Q. Chen (Triuridaceae) is described and Island, China. It differs from the closely related S. arfakiana Becc. by its longer leaves to ca. 2 mm (vs. 3-6 mm (vs. typically 7-9 mm), three stamens (vs. two or three). 2-celled anthers (vs. 4-celled). and a filiform style (vs. awl-shaped) that far exceeds the Key words: China, Hainan, IUCN Red List, van de Kamer & Weustenfeld, 1998), with about 50 New World tropics (Meerendonk, 1984; Maas & Riibsamen. 1986; Mabberley. 2008; Guo & Cheek, 2010). the genus by the following features: small, achloro- and * actinomorphic flowers, with (four to) six (to 10) perianth segments basally connate. The male flowers have two to six stamens, with 1- to 4-celled anthers flowers consist of ca. 10 to 80 ovaries, each with a persistent stamens, with 1-celled anthers and ca. 10 and are not easily discerned in the field. Only five Wu et al.. 2000: Ye. 2003: Hsieh et al.. 2003; Zhuang et al., 2004; Guo & Cheek, 2010): S. arfakiana Becc.. S. maculata Miers. S. ramosa (). Chen, sp." nov. " TYPE: China. Hainan: Jianfengling Natl. Nature Reserve, under trop¬ ical rain forest, 18°44'N, 108°53'E, 810 m, 12 Aug. 2007, Han Xu & H. Q. Chen JFL00891 (holotype, CANT). Figure 1. glabrous; stems erect, slender, with 1 or 2 branches ca. 8-11 cm, 0.3-0. 5 mm diarn. Leaves alternate, flowers; pedicels 3-6 mm, 0. 1-0.2 mm diarn. , straight: bracts lanceolate. 1.2-2. 1 mm; tepals 6, Novon 21: 154-157. Published on 7 April 2011. doi: 10.3417/2009016 Volume 21, Number 1 Xu et al. 155 Figure 1. Sciaphila filling™* Han Xu, Y. D. Li & H. Q. Chen. -A. Single fertile individuah -B. Sealelike feaf. -C. Lu from holotype Han Xu & H. Q. Chen JFL00891 (CANT). ? ? 7 7 156 IUCN Red List category. Hainan Island is characterized by a high level of endemism. The 157 Acknowledgment of Reviewers 159 Hong Wang Hui Wang Volume 21, Number 1, pp. 1-160 of NOVON was published on 7 April 2011. www. mbgpr ess . info CONTENTS ^jjlk Missouri 1— c /a4- o ~t s~\ o I Botanical Garden A Journal for Botanical Nomenclature VOLUME 21 NUMBER 2 2011 Volume 21, Number 2 June 201 1 Novon, A Journal for Botanical Nomenclature from the Missouri Botanical Garden The mission of the Missouri Botanical Garden is to discover and share knowledge about plants and their environment, in order to preserve and enrich life. Novon publishes short articles whose primary purpose is the establishment of nomencla¬ ture in vascular plants and bryophytes. All articles are peer-reviewed by qualified, indepen¬ dent reviewers. Manuscripts must fully state and justify the reasons for proposing nova. These may include detailed comparisons with similar taxa, short keys to similar taxa, illustrations to similar taxa, and mechanical nomenclature reasons, among others. Manuscripts whose primary purpose is other than establishment of new nomenclature, which usually are longer manuscripts, cannot be accepted for review. These include reviews, revisions, monographs, or other papers that incidentally include nova. Manuscripts must follow the guidelines in the Checklist for Authors. The Checklist may be downloaded from the Garden’s web site, www.mbgpress.info, or authors may contact the man¬ aging editor to request a copy. Novon will not knowingly accept manuscripts that have been simultaneously submitted to other journals for consideration or previously published in some form elsewhere. Victoria C. Hollowell Editor Beth Parada Managing Editor Allison M. Brock Associate Editor Tammy Charron Associate Editor Cirri R. Moran Press Coordinator Ihsan A. Al-Shehbaz Consulting Editor Carmen Ulloa Ulloa Consulting Editor George Yatskievych Consulting Editor Kanchi N. Gandhi Nomenclature Consultant Nicholas J. Turland Nomenclature Consultant Roy E. Gereau Latin Editor Novon is included in the subscription price of the Annals of the Missouri Botanical Garden. Price for 2011: $180 per year U.S.A.; $190 Canada and Mexico; $215 all other countries. Four issues per volume. Postal address: Subscriptions: Annals of the Missouri Botanical Garden, P.0. Box 1897, Law¬ rence, KS 66044-8897; Manuscript submissions: Novon, Missouri Botanical Garden, P.O. Box 299, St. Louis, MO 63166-0299. Web site: http://www.mbgpress.info © Missouri Botanical Garden 2011 © Novon is printed on paper that meets the re¬ quirements of ANSI/NISO Z39.48-1992. Novon (ISSN 1055-3177) is published quarterly by the Missouri Botanical Garden, 2345 Tower Grove Avenue, St. Louis, MO 63110. Periodicals postage paid at St. Louis, MO and additional mail¬ ing offices (USPS #006-777). POSTMASTER: Please send address changes to Annals of the Missouri Botanical Garden, P.O. Box 1897, Lawrence, KS 66044-8897. Information on the contents of Novon is present¬ ed in SciSearch®, Research Alert®, Current Con¬ tents®^ lture, Biology IE r n ental Sciences, APT Online, the CAB Abstract/Global Health database, and the ISI® database. The full-text of Novon is available online through BioOne™ (http://www.bioone.org). Two New Species of Uvaria (Annonaceae) from Borneo, with a New Nomenclatural Combination A. M. Achala S. Attanayake School of Biological Sciences, The University of Hong Kong, Pokfulam Roach Hong Kong, China. Current address: Deputy Director, National Herbarium, Department of National Botanic Gardens, Peradeniya, Sri Lanka, achala.attanayake@gmail.com Ian M. Turner Research Associate, Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, United Kingdom; Research Associate, Herbarium, Singapore Botanic Gardens, Cluny Road, Singapore 259569, Singapore, j_trop_ecol@yahoo.co.uk Richard M. K, Saunders School of Biological Sciences, The University of Hong Kong, Pokfulam Road, Hong Kong, China. Author for correspondence: saunders@hkucc.hku.hk Abstract. Two new species of Uvaria L. (Annona¬ ceae), U. beccarii Attanayake, I. M. Turner & R. M. K. Saunders and U. curvistipitata Attanayake, I. M. Turner & R. M. K. Saunders, are described and ing a legitimate name in Uvaria for the species. The in the Annonaceae (KeBler, 1993), with ca. 210 of diversity in continental Southeast Asia and Malesia (Meade. 2005). All species of Uvaria are woody twining branchlets. The genus is characterized by the presence of stellate hairs (throughout the plant), generally arranged in two lateral rows (Meade, 2005; f. & Thomson. EUipeiopsis R. E. Fr.. Rauivenhoffia in phylogenies based on chloroplast DNA sequence data (Zhou et al.. 2009. 2010). The diagnostic Peninsular Malaysia (Sinclair, 1955) and continental Asia north of the Isthmus of Kra (Meade, 2000) — no indicate that there are over 30 species in the region, here, associated with the recent transfer of Cyatho- stemma species to Uvaria (Zhou et al., 2009). 1. Uvaria beccarii Attanayake, I. M. Turner & R. M. K. Saunders, sp. nov. TYPE: Malaysia. Sarawak: Mt. Matang, 1863-1865 [1865, fide van Steenis- doi: 10.3417/2009123 Novon 21: 161-168. Published on 27 June 2011. 162 2011 164 Novon Volume 21, Number 2 2011 Uvaria (Ai et al. 2011 ff of E, FI, K, KEP, L, SAN, SAR, a A New Species of Pera (Euphorbiaceae) from Amazonian Brazil Narcisio Costa Bi.gio Universidade Federal de Rondonia, Depto. Biologia, Campus Jose Ribeiro Filho, BR 364, Km 9.5, 76800-000, Porto Velho, Rondonia, Brazil, narcisio@gmail.com Ricardo de S. Secco Museu Paraense Emilio Goeldi, Depto. Rolan iea. C.P. 399, 66040-170, Belem, Para, Brazil. rsecco@museu-goekli.br the Brazilian Amazon, Pern eiteniorum Bigio & closely related to P. tomentosa (Benth.) Mull. Arg., P. late flower with a lagenifonn (flask-shaped) ovary. semelhante a P. tomentosa (Benth.) Mull. Arg., P. Johnst., mas separa-se por ser a unica deste grupo a das folhas com tricomas estrelados e estrelados estaminadas glabros ou raramente com tricomas tamente oblanceolado. Discussao sobre as afinidades Amazonia brasileira. e apresentada. 1997), occurring from Cuba and Central America to exemplified by P. distichophylla (Mart.) Bail! and /’. uses: P. glabrata (Schott) Baill., which is used to al„ 1992). Gillespie and Armbruster (1997), treating five and Hoffmann (1919). Traditionally, Pera has been studies using DNA sequence data have shown that al., 2005; Wurdack & Davis, 2009). In order to the Peraceae (Stevens. 2001 onward; Souza & Lorenzi, 2008; Wurdack & Davis, 2009). However, APG III (2009) still does not recognize Peraceae as a In a revision of the Pera species occurring in the including types from IAN. INPA. MG. R. RB. SP. UB, and UFMT, and photographs of the types from F, G, IAN, K, L, NY, and P, we found diverse doi: 10.3417/2009089 Novon 21: 169-173. Published on 27 June 2011. 170 Novon 171 2011 rs«, and disc a do Garcas, 254 k 8499 (holotype, SP). Fi^.re 1. 172 173 2011 to Lynn Gillespie, Hollowell for their Typification of Neotropical Species of Staurogyne (Acanthaceae) C.P. 199, Volume 21, Number 2 2011 Braz & Monteiro Typification of Neotropical Staurogyne 175 year, Nees (1847b) included material from Martius and four numbered specimens by Riedel (5, 10, 348, Baptistae. mense Aprili”), Riedel 10 (“Mariana, Julio 1824”), and Riedel 348 (“bracteis lutei floribus publication (Nees, 1847b: 72) and can be inferred to represent the syntypes (Pohl 3151, W; Vauthier 182, G; Riedel 348, LE). Among the syntypes, only Pohl’s collection corresponds to the distinct taxon that we leaves 9.5-19 cm long, petiole 0.6-1. 8 cm, and designate Riedel 348 as lectotype because of its orbicular and colored bracts, as well as the short Boiler 5(8 1: 615. 1897. TYPE: Bolivia. [Santa Cruz: Jose Miguel de Velasco], “200 m alt.,” here, US 702151; duplicates, GH not seen, NY 278273). Velasco orient. Alt! 200 m (0. Kuntze flor. Julio),” this accession was destroyed in World War II. the sheet was photographed and is now at the Field Museum (F 043983). Because no holotvpe was three duplicates are known from GH, NY, and US. petioles 1.1-2. 3 cm, and corolla 32-44 mm. Boi--icr 5(8): 611. 1897. TYPE: Brazil. Santa Catarina: Blumenau. Aug. 1884. C. A. W. 45230). ogyiie in the Americas, we were unable to locate the 1897: 644, “S“ Catarina, prope Blumenau in silva ad Heinrich Schenck traveled in Brazil in 1886-1887, totally destroyed during World War II (Stafleu & attributes cited by Lindau (1897) in the description same general locality as the original type. PI. 2: 497. 1891. Basionym: Ebennaier a elegans Nees, FI. Bras. (Martius) 9: 17-18. 1847. TYPE: Brazil. Minas Gerais: “ad Vieira do Matto,” s.d., J. B. E. Pohl 3151 (lectotype, designated here, W; duplicates, BR 840386, GZU 251331). PI. 2: 497. 1891. Basionym: Ebennaiem hirsuta Nees. FI. Bras. (Martius) 9: 18. 1847. TYPE: Brasil. Minas Gerais: “Tejuco,” 1883, Vauthier GZU 251310). Pohl (“ad Vieira do Matto”), Vauthier (“ad Viliam by the hirsute-tomentose stem, the lanceolate leaf other characters. The collections Schiich s.n. and 176 Novon (Nees, 1947a), all collected in the same locality Schiich”). Later, Nees numbered Vauthier 18.3 and Museum Wien (W), in addition to the collection Gen. PI. 2: 497.' 1891. Basionym: Ebermaiera mandioccana Nees, PI. Asiat. Rar. (Wallich) 3: 80. 1832. TYPE: Brazil. Rio de Janeiro, s.d., L. Brazil, dating front 1822 to 1823, were distributed in many herbaria. The cited holotype (Nees, 1832: 80, a. 1822, cum floribus fructuque maturo legit genus in the 20th century (Leonard, 1951, 1958; Wasshausen & Smith, 1969; Kanteyama, 1991). Nees (1847a: 16, 1847b: 71) later mentioned the collec¬ tions of Pohl 4740, Pohl s.n. (“prope Rio de Janeiro”), and Riedel 797 (“in silvis Macahe”), in of Riedel (Riedel ^797, LE, “in silvis Macahe”) both consists of plentiful material and conforms to the TYPE: Brazil. Minas Gerais. Sep. 1841, G. Gardner 5129 (lectotype. designated here. W: duplicates. BM, E, FI, K, P, R). minarum (Nees. 1847a). the collections Martins 943, Ackemiann s.n., Schiich s.n., Gardner 5129, Sellow s.n., and Riedel s.n. were cited and are the herbaria and collection numbers as Martius 94.3 (M). Sellow 151, 210. 25.3 (B). and Riedel 6. 467 25.3 (B, duplicate at K) represent S. minarum based descriptions of Nees (1847a. 1847b) but has not been (BM, E, FI, K, P, and R) that consist of copious PI. 2: 497. 1891. Basionym: Ebermaiera repens Nees, FI. Bras. (Martius) 9: 20. 1847. TYPE: Brazil. Mato Grosso: Mun. Cuiaba, June 1827, L. Riedel 1087 (lectotype, designated here, LE). other in “Serra da Chapada.” Nees (1847b) later LE) for this species, two of which (Riedel 1 and 1087) leaves (0.9-5. 3 X 0.3-0. 7 cm). Both of these elliptic leaves (0.9-2. 3 X 0 3-1.2 cm). The specimen Gen. PI. 2: 497. 1891. Basionym: Ebermaiera riedeliana Nees. FI. Bras. (Martius) 9: 18-19. 1847. TYPE: Brazil. Rio de Janeiro, s.d., L. 177 2011 A New Species of Cohniella (Orchidaceae, Cymbideae, Oncidiinae) from Amazonian Venezuela William Cetzal lx El Colegio de la Frontera Sur, Unidad Chetumal, Av. del Centenario, Km 5.5, Chetumal 77000, Quintana Roo, Mexico, rolito22@hotmail.com German Carnevali Femdndez-Concha Herbario CICY, Centro de Investigacion Cientffica de Yucatan, A.C., Apartado Postal 87, Cordemex, Merida 97310, Yucatan, Mexico. carneval@cicy.mx Venezuela. The new species is illustrated and its that emerge directly from the labellum disk, with the proximal half. In contrast, the callus in C. cebolleta is Reslimen. Una especie nueva fue encontrada du¬ rante nuestro trabajo monografico en el genero cebolleta (Jacq.) Christenson del norte de Venezuela The members of this orchid genus can be easily flowers (Carnevali et al., 2010). Characters used to shape and number of teeth of the callus of the lip, and Additionally, we have used diagnostic elements for orientation of the plants (e.g., pendent or erect), leaf & Carnevali, 2010). tmm Poepp. & Endl.. has been controversial. As Terrazas. 2001: Sosa et al.. 2001: Williams et aL 2001a, 2001b; Chase, 2009) who have treated and floral traits. The rationale behind this narrower elsewhere (Pupulin & Carnevali. 2005: Cetzal et al., 2008; Carnevali et al., 2009, 2010). Our Novon 21: 178-181. Published on 27 June 2011. doi: 10.3417/2009063 2011 Novon 2011 Venezuela. The labels of the specimens from the a (IUCN, 2001). Although the r, A New Name for Papaver pseudo-orientale (Papaveraceae) 2011. 21: 182. doi: 10.3417/2009134 Convolvulaceae Neotropicae Novae vel Minus Cognitae, I. Bonamia jiviorum , a New Species from the Venezuelan Guayana Jose Ramon Grande Allende Postgrado en Botanica, Facultad de Ciencias, Universidad Central de Venezuela, Calle Suapure, Colinas de Bello Monte, Apartado 47114, Caracas, Distrito Capital, Venezuela. jose.r.grande@gmail.com Abstract. The new species Bonamia jiviorum J. R. Reslimen. Se describe e ilustra Bonamia jiviomm J. R. Grande ( Convolvuloideae, Cresseae) del norocci- irregulannente estriada. con alas sobre las aristas ciudad de Puerto Ayacucho. donde se encuentra long or more (Myint & Ward. 1968). Species of al, 2002, 2003), an excellent study object " in Austin, 1982, 1994; Austin & Staples, 1985; Johnson. 1987; McDonald, 1987; Gentry, 1988 [1989]; Breteler, 1992; Deroin, 1992 [1993], 1998, 2004), as well as other important changes in the doi: 10.3417/2008097 Novon 21: 183-186. Published on 27 June 2011. TYPE: Volume 21, Number 2 2011 Grande Allende Bonamia jiviorum (Convolvulaceae) 185 A New Species of Aspidistra (Ruscaceae) from Guizhou, China Novon He et al. 201 1 ’ Aspidistra (Ruscaceae) from China Valid Publication of Asarum longirhizomat.osum (Aristolochiaceae) Nan Jiang Institute of Subtropical Crops, Zhejiang Academy of Agricultural Sciences, Wenzhou 325005, Zhejiang, People's Republic of China; and Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650204, People's Republic of China, jiangnan@mail.kib.ac.cn Xiao-Ming Peng Academy of Forestry Inventory and Planning, Jiangxi, Nanchang 330046, People's Republic of China Wen-Bin Yu Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650204, People's Republic of China Liang and C. S. Yang in 1975, was invalidly collections were cited as type. The name is validated here by designating the collection Chao-Liang Zhang 002 (IBK 00190377) as the holotype. The genus Asarum L. s.l. (Aristolochiaceae), with America and Europe (Kelly, 1998; Huang et al., (Liang, 1975). This name has been widely used as it Sinicae (Cheng & Yang. 1988: 192). and subse¬ quently in the English edition. Flora of China (Huang et al., 2003: 256). However, the name was not validly “typus” in the original description (Liang, 1975: 21). Liang Zhang 002, were both separately and (McNeill et al., 2006) for names published on or after Furthermore, upon contact to CMMI. the collection Liang & C. S. Yang. The spelling of the epithet is dation 60G. Sin. 13(2): 21, pi. 1, fig. 2, pi. 2, fig. 4-10. 1975 Darning Mtn.. 1972. Chao-Lian Zhang 002 (holotype, IBK 00190377; isotypes, IBK 00190376, 00190378). Novon 21: 190-191. Published on 27 June 2011. doi: 10.3417/2009067 2011 Jatropha mirandana (Euphorbiaceae), Especie Nueva de la Cuenca Oriental del Rio Balsas de los Estados de Guerrero y Puebla, Mexico 2011 194 Novon — G. Semilla. A-C tornado del holotipo Jimenez & Vega 2009-128 (FCME); D, E de Jimenez & Vega 2009-126 (FCME); F, G de Contreras 981 (FCME). 2011 Balsas (Sapindaceae), Genero Nuevo de la Cuenca del Rio Balsas en el Estado de Guerrero, Mexico Volume 21, Number 2 2011 Jimenez Ramirez et al. Balsas (Sapindaceae) de Mexico 197 >s de 3 a 9 flores, con 2 z; Novon — Q. Semilla, vista la 18 (FCME); L-Q, m; las e; i & Jimenez 1314 j 1319 (FCME); R de Cruz 7325 (FCME). Las estructuras B-K e 2011 resto de los , la clave de males (3.7-)4.5-6.1(-7.2) X (2-)2.6-3.3(-3.8) Deciolulo (0.2-)0. 4-0.8 cm; los folmlos temii- (4.8-)7.6-8.4(— 9) X (3-)4-4.8(-5.8) cm. Paratipos. MEXICO. Guerrero: lutla, 6.66 km al SE, 1555 m, 1 die. 2.5-3 .'2007>(frL)!RC^:S5 Three New Distichous-bracted Species of Calathea (Marantaceae) from Panama the family, with 14 in the (1976: 312) it of ca. 160%. Six of tl Marantaceae is the region near Portobelo (Colon 2011. 2011 1, 2-3 X 2. 8-4. 4 c: . 1. 6-2.1 X 1. 1-1.6 cm. 0. 6-1.1 cm wide cent pale cream-yellow. (1.5-)1.8-2.1 X (1.1-) 7 cm; hracteoles 1 per flower pair, indurate X14. 12-16 X 3.5-5 mm; rounded, cupped, yellow or pink-purple. 11-13 X 9- 204 Novon the ovate leaves with length :width ratios of (1.7-) length:width ratios (more than 1.7:1 vs. less than 2011 ( I ll I iquensis H. Kenn., sp. nov. TYPE: Panama. Chiriquf: vie. of Gualaea ea. 8.5 mi. from Planes de Homito, La Fortuna on rd. to dam site, near stream. 4400 ft.. 10 July 1980. T. MO. UBC). t. hairs ca. 0.1 mm 2011 BSBBS9 Volume 21, Number 2 2011 210 2011 Elat.ostema oppositum (Urticaceae), a New Species from Yunnan, China State Key Laboratory of S 21: 212-215. 2011. Volume 21, Number 2 2011 Lin et al. 214 Lin et al. 2011 A New Species of Astragalus (Leguminosae) from Northwestern Xinjiang, China types with two kinds of at -25 cm; petiole 7-10 cm, t, with leaflets in 5 to 9 pairs; 15(— 20) X (4-)6-8(— 10) mm, acute to obtuse, often sparse black hairs; racemes oblong capitate, 2.5-3.5 X 1.5-2 cm, each raceme congested, with 12 to 15 flowers. Calyx in anthesis tubular. 15-18(-20) mm. 19-21 mm, with limbs narrowly oblong, 6-7 mm. asymmetric-ally emarginate at apex, claw 1.5X as long as limb, auricle ca. 1 mm; keels slightly shorter than wings, 17-19 mm, with limbs obliquely elliptic 5-7 mm. claw 1.5X as long as limb, auricle 0.5-1 mm; 2011. Volume 21, Number 2 2011 Liu et al. A Multivariate Study of Solidago subsect. Junceae and a New Species in South America (Asteraceae: Astereae) Rita M. Lopez Laphitz Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada. Current address: Departamento de Botanica, INIBIOMA (Universidad del Comahue-CONICET), Quintal 1250, CP 8400, San Carlos de Bariloche, Argentina Yimfei Ma Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada. Current address: MD Program, Faculty of Medicine, University of Toronto, 1 Kings College Circle, Toronto, Ontario M5S 1A8, Canada John C. Semple Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada. Author for correspondence: jcsemple@sciborg.uwaterloo.ca Abstract. A new South American species of tion Junceae (Rydb.) G. L. Nesom. Using multivariate analyses on a matrix of 50 characteristics for 79 S. missouriensis-Uke taxon was tested and determined t Small. Therefore, a new South Triplinerviae (Torr. & A. Gray) G. L. Nesom were DC. (Lopez Laphitz & Semple, 2011). In addition, the riensis Nutt. Consequently, these specimens were excluded from the analysis of Solidago subsect. Mexico (Semple & Cook, 2006): S. gattingeri Chapm. A. Gray from the mountains of the Great Basin, the northern Baja California Norte. Solidago subsect. characteristically with small linear leaves on very (Semple & Cook, 2006). Strigulose stem pubescence Although Brouillet et al. (2009) were successful in nuclear ribosomal ITS data. Schilling et al. (2008) levels of variation in ITS and ETS sequences. Until analysis to assess the distinctiveness of the South doi: 10.3417/2010088 Novon 21: 219-225. Published on 27 June 2011. Volume 21, Number 2 2011 Lopez Laphitz et al. 221 Solidago subsect. Junceae (Asteraceae) pinetorum. and 7% to S. gattingeri, Of the 11 variant scores were plotted on the first and second assigned a posteriori to that taxon with Geisser putative taxa studied (Fig. 1). The eigenvalues for the was assigned corrected to the species with only 45% the variation, the second (0.573) accounted for probability. Three of the four misclassified specimens 14.4%). and the third (0.249) accounted for an were placed a posteriori in S. pinetorum (50%) — 60%) additional 6.3%). probabilities), and one was classified as S. misson- The results of analysis of Solidago argentinensis, S. riensis (53%). chilensis, and S. microglossa confirmed that the three juncea, 10 were assigned a posteriori to that taxon; the 15 specimens treated a priori as S. argentinensis between 71%) and 100%). with six specimens between probability for all. including the single Chilean 49%) and 67%). The remaining three specimens were specimen, which resembled S. chilensis in general between 2%) and 40 %. Of 26 specimens assigned a priori to S. missouriensis, 15 were placed into that Discussion probabilities between 49%) and 56%). Three of these South American species is recognized in Solidago Among the North American specimens classified a species (S. gattingeri, S. jnncea, S. missouriensis, and tests for 222 Novon (. Pedersen 14486. MO) was from Chubut Province, the species. It was placed a posteriori with 100% microglossa. the three native species of Solidago in sions about their status can be reached. (2009) and Lopez Laphitz and Semple (2011). This can be explained by the morphological similarities Ariza Espinar. 2003). In addition, the overlapping missouriensis on the high plains in North America. usually reliable character such as stem height is Solidago. " ; ^ America has not been studied in detail, the chance (Alan Graham, e- Table 4 2011 224 Novon Rinconada y Junin de los Andes, A. L. Cabrera & J. Crisci 19130 (LP). Rio Negro: Destacamento Militar (Parque Nacional Nalmel Huapi). R. De Barba 301 (AA); Puerto Panuelo, region of Lago Nalmel Huapi, /. Rafael Cordini 121 (LP); camino al Tronador, C. Castagnet 74 (LP); San Carlos de Bariloche, A. Burkart 6586 (LP), A. Corte 291 2011 New Species and Notes in the Genus Dioclea s.l. (Fabaceae, subtribe Diocleinae) Richard H. Maxwell Indiana University Southeast Herbarium (JEF), 4201 Grant Line Road, New Albany, Indiana 47150, U.S.A. rmaxwell@ius.edu Abstract. Seven new species are described in R. H. Maxwell is described from Colombia and the new series Virgatae R. H. Maxwell is created, Bentham are raised to subgenera status, as the new H. Maxwell and Dioclea subg. Platylobium (Benth.) R. H. Maxwell. Within subgenus Platylobium , D. species are described: D. apiculata R. H. Maxvell Colombia, D. haughtii R. H. Maxwell from Colombia Bentham (1837) divided Dioclea Kunth (Fabaceae, Diocleinae) into section Eudioclea Benth. and section Bentham (1859) later added one new species, and third section, section Platylobium Benth. To separate his three sections, Bentham (1837, 1839) used tions Dioclea subg. Platylobium (Benth.) R. H. H. Maxwell are established. Amshoff (1939) named a represent some originally cited in my doctoral dissertation (Maxwell, 1969). Lackey (1981), in his key. split Dioclea into two more, two articles (Maxwell & Taylor. 2003: Queiroz et al., 2003) show the existence of separate clades of tional taxonomic units [OTUs]) were selected in subgenera and the sections or series in Maxwell (1969). Maxwell and Taylor (2003) stated that the of species in Dioclea. Queiroz et al. (2003) found a clade. with Dioclea species in the latter all within lobi-um clade. I assert that the Dioclea species in the Cymbosema-Dioclea clade (Queiroz et al., 2003) and Dioclea subg. Dioclea (Maxwell & Taylor, 2003) Varela et al. (2004: 61), using spacer sequences Taylor. Queiroz. and Maxwell have agreed to groups within Dioclea s.l. DNA analysis of Dioclea lobi-um Sm. (Smith, 1793), and if Dioclea subg. rejlexa Hook. f. (= D. hexandra (Roxb.) Mabb.), Novon 21: 226-243. Published on 27 June 2011. doi: 10.3417/2002014 2011 228 Novon 2011 i ca. 7 X 2. 5-3. 5 i 2-2.5(-3) ■. 8-14.5 X 4. 5-7. 5 230 Novon Volume 21, Number 2 2011 Maxwell 231 Dioclea s.l. (Fabaceae, subtribe Diocleinae) slightly cordate, adaxial lamina surface glabrate 30-65 cm. terete, with short, ascending, curly, canescent pubescence, florate ca. 1/2 the length; ral bracts acute or ovate, ca. 2.5 mm. glabrate, persistent; bracteoles ovate, ca. 6-10 X 6-8 mm. ca. 18 X 19 mm. claw ca. 6 mm; wings somewhat shallowly serrate, occasionally entire; stamens 10. glabrous, vexillary filament free at the base ca. 4 mm. basal disc collar, ca. 8-ovulate, with whitish 2-3 X 0.4 cm. eventually dehiscent, with stiff, erect. X 8.5 X 2.5 mm. hilum ca. 14 mm. encircling nearly cies is found in swampy forests, woods, wooded cliffs. Colombia, west of the Cordillera Occidental, at low has the smallest bracteoles (ca. 3 X 3 mm) and flower pedicels (ca. 3 mm long). Other species in series southern Brazil, and Paraguay; D. fimbriata along roseum Benth. into Dioclea [= Dioclea rosea (Benth.) N. Zamora]. Queiroz et al. (2003: 317) indicated that present study (0.17) and the relationship of Cymbo- its taxonomic placement.” Placement of D. macran- tha next to C. roseum occurred in Maxwell and philous (Queiroz et ah. 2003: 305. 317)! Varela et al. parallel to the upper suture, ca. 4.4-6 X 1.8-2. 2 X O. 4-0. 7 cm. with a downcurved beak, and two to five seeds (Maxwell. 1970. 1999; Kirkbride et al.. 2003). Paratypes. COLOMBIA. Choco: Rio San Juan, J. Cuatrecasas 21380 (F); Docordo, N mouth of Rio San Juan, A. Gentry, E. Zardini, M. Monsalve & J. D. Caicedo 53845 (JEF, MO). Valle del Cauca: Dept, del Valle, along Rio Anchicaya, J. Cuatrecasas 15257 (F, US); del Valle, Pacific Volume 21, Number 2 2011 mil 2011 rarely 10 perfect uniform: pistils with the s usually proximally swollen, ovary disc dentate. F indistinct, (1 to)2- to 9(to Ill-seeded; seeds various. [1990a: 581]). round to slightly oblique: primary lateral veins in 10 to 14 pairs; petioles ca. 8-14 cm, rachis ca. 1/4 the , 13-15 X ca. 10 n 236 Novon 2011 238 Novon 2011 12 X 8 cm, lateral leaflets ca. 12 X 7.5 cm, adaxial ca. 10-12 mm, then geniculate ca. 7-9 mm. style distally bent back, glabrous ca. 2 mm: ovary ca. 5 mm, villous with canescent or fuscous hairs 1.5-2 2- to 4(to 5)-seeded; 4.5 X 2.4-4 X 4 1889,” sm. (holotype. 240 Novon & Gonzalez 16,219 (MO). 2011 15 X 6 X 1.4 X 6 X 1.4 cm; Paratypes. ECUADOR. Loja: betw. Rfo Camba & Loja city, July 1876, E. Andre 4196 (K); Los Rios, Jauneche forest, fruits black, in swamp, 24 Jan. 1981 (fr.), A. Gentry, vine along Rio Baba, S of St Dodson & L. Thien 1200 (M :t for the Ph.D. by John D. Dwyer (1915-2005) 242 Novon 2011 Cestrum sotonunezii, a New Species of Solanaceae from Central Mexico Juan Carlos Montero-Castro Facultad de Biologfa, Universidad Michoacana de San Nicolas de Hidalgo, Ciudad Universitaria, Edificio B2, 31'1 pi-o. Morelia. 50030 Miclioacan. Mexico. ce-lmm2()().'5@yahoo.com.mx Lindl., C. oblongifolium Schltdl., and C. thyrsoideum Kunth, all of which also occur in central Mexico. The sotonunezii Mont. -Castro (Solanaceae) proveniente del centro de Mexico. Las colecciones de la nueva flcwescens Greenm., C. aurantiaciun Lindl., C. caracterfsticas. especialmente en los estamhres. Se diferenciar especies morfologicamente similares del centro de Mexico (Guerrero, Michoacan). Mexico, Michoacan, Solanaceae. (1998). The new species C. sotonunezii Mont. -Castro, (Endl.) Schltdl., has highly attractive, club-shaped to classify species in this section (Francey, 1935). Novon 21: 244-248. Published on 27 June 2011. 2006). aurantiacum Lindl., C. oblongifolium Schltdl., and C. as well as by the length of its corolla lobes. The and allied species in central Mexico that are characterized by having 5- to 25-flowered axillary ing ca. 2 mm above an entire bulge; corolla lobes < ' below or directly from a bilobed bulge; corolla lobes 1.3-2.8 m doi: 10.3417/2005040 2011 Leonardo Bravo], Filo de Caballo, 2190 m, 22 May 1986, J. C. Soto Nunez 12678 (holotype, MEXU; isotypes, MO, NY, US). Figure 1. Shrubs 1-3 m high, flowering from the end of dry ovate to widely ovate (young leaves elliptic), 3.7-10 X 2-5.8 cm, cuneate at base, acuminate at apex, lateral s near Filo de Caballo, ( Basin. Although the areas of occupancy are very tight calyx at the base of the corolla, and the calyx length is less than one third of the corolla length. In contrast, the calyx of C. thyrsoideum is loosely placed 246 Novon Volume 21, Number 2 2011 247 Neotypification of Hybanthus longistylus (Yiolaceae) A New Monotypic Genus, Ananthura , from Tropical Africa (Asteraceae, Vernonieae) Harold Robinson Department of Botany, MRC 166, National Museum of Natural History, P.0. Box 37012, Smithsonian Institution, Washington, D.C., 20013-7012, U.S.A. Robinsoh@si.edu John J. Skvaiia Department of Botany and Microbiology, and Oklahoma Biological Survey, University of Oklahoma, Norman, Oklahoma, 73019-6131, U.S.A. jskvarla@ou.edu Abstract. A new genus. Ananthura H. Rob. & Skvarla (Asteraceae, Vernonieae), is described for Vernonia pteropoda Oliv. & Hiern in Oliv. of Central Africa. The genus has broad median shields on the The name V. urophylla Muschl. is lectotypified. species (Robinson, 1999a) and most of the tropical American species (Robinson, 1999b) that were alternative dispositions continues here with the shields on the involucral bracts place the genus in the Methods in Hoyer’s solution (Anderson, 1954). Preparation of consisted of acetolysis (Erdtman. 1960) followed by of Oklahoma with a JEOL 880 SEM (JEOL Ltd., Results and Discussion Jeifrey (1988: 213-215) placed Vernonia pteropoda inner phyllaries and ± 10-ribbed achenes.” Jeffrey some in the subtribe Gymnantheminae (Keeley & defined by Jeffrey (1988), V. pteropoda seems to stand petiole that has a tapered narrow wing reaching subtribe in Robinson (1999a, 2007) and the narrower sense of Keeley and Robinson (2009). Placement of Vernonia, pteropoda in the genus doi: 10.3417/2010062 Novon 21: 251-255. Published on 27 June 2011. 252 Novon Volume 21, Number 2 2011 2011 Fagopyrum wenchuanen.se and Fagopyrum qiangcai , Two New Species of Polygonaceae from Sichuan, China Ji-Rong Shao,1 Mei-Liang Zhou Xue-Mei Zhu,2* De-Zhou Wang,1 and Da-Quan Bai 1 'School of Life Sciences, Sichuan Agricultural University, Yaan, Sichuan, (>25014, People's Republic of China 2School of Resources and Environment, Sichuan Agricultural University, Yaan, Sichuan, 625014, People's Republic of China *Authors for correspondence: zhml39@yahoo.cn, shaojr007@163.com Province, China, are described and illustrated. lentum Moench based on its leafy base, triangular leaves, and terminal or axillary racemes. The new differs in its caespitose habit, the plants often with variable in length, and the capsules are ellipsoid- triangular or broadly ovoid-triangular, 3-3.5 X 2.5-3 Key words: China, Fagopyrum , IUCN Red List, Fagopymm Mill. (Polygonaceae). It is an annual species are found (Li, 1998; Ohnishi, 1998). In 4500 m (Wang, 1989; Ohnishi & Yasui, 1998). Ohnishi (1998) reported that the cultivated common China to the Himalayas. In July 2006, we found two as F. qiangcai D. Q. Bai and F. wenchuanense J. R. Materials and Methods Agricultural University. Selected seeds were soaked distilled water, and hydrolyzed in 1 mol/L HC1 at and Chen (1985); the chromatid classification Taxonomic Results 1. I i | m u in «|i ii ii D. Q. Bai, sp. nov. TYPE: growth of grass on slopes, 1580 m, 16 July 2006, J. R. Shao & D. Q. Bai Ghassot 4396 (holotype, SAU). Figure 1. Novon 21: 256-261. Published on 27 June 2011. doi: 10.3417/2009107 Volume 21, Number 2 Shao et al. 257 Volume 21, Number 2 2011 Shao et al. Fagopyrum (Polygonaceae) from China 259 ** * r % * ♦ r a: 4. 4 * * * "tjs W hr u Ri n n u u M tttt D w R n II M If U K « RUN 0» (I It n bi n ideograms. A, C from Bai Ghassot 4472 ; B, D from Bai i (all at SAU). w nr it u It X| IF U cell chromosomes. — B. F. gracihpe. s (Hemsl.) Dammer ex Diels, f 4491 ; E, G from Bai Ghassot 4483 ; F, H from Bar Ghassot 4475 260 Novon et al. 2011 (Polygonaceae) from China Hshh'g Co!, Blij^g [in Chinese]^ ailemK JOUmal ^ A New Species of Epimedium (Berberidaceae) with 24 Chromosomes from Guizhou, China Mao Yin Sheng1'2 and Xing Jun Tian1* 'School of Life Science, Nanjing University, Hankou Road No. 22, Nanjing, 210093, People's Republic of China institute of South China Karst, Guizhou Normal University, Baoshanbei Road No. 116, Guiyang, 550001, People's Republic of China * Author for correspondence: tianxj@nju.edu.cn Abstract. The new species Epimedium yinjian- gense M. Y. Sheng & X. J. Tian (Berberidaceae) is to E. acuminatum Franch. but differs in its Red List, tetraploid. ease (Guo & Xiao. 1999). as well as to strengthen worldwide, distributed from the Mediterranean re- (Ying, 2002). The center of diversity for the genus is in China (Ying, 2002). and 52 taxa of Epimedium were reported as native to China (Guo et al., 2008). (Stearn. 2002). Species native to China are grouped morphologically, with distinct inner sepals and a including the Flora of China ( Ying et al., 2011), as sp." nov." TYPE: China. Guizhou: Tongren, 1300 mS 28°02'E, 108°42'15"N, 5 Mar. 2004, M. Y. Sheng 040308 (holotype, GNUG; isotype, N). Figures 1, 2A, B. Perennial herbs. 40-60 cm tall: rhizomes rigid. ovate to narrowly ovate. 4. 1-5.6 X 2. 1-2. 7 cm, cordate, the terminal leaflet with equal, rounded 4, obovate, 4-5.5 X 3-4 mm, caducous; inner sepals 20(— 25) mm, white or light "mulberry-purple; stamens Novon 21: 262-265. Published on 27 June 2011. doi: 10.3417/2009081 Volume 21, Number 2 2011 Sheng & Tian Epimedium (Berberidaceae) from China 263 a 1000 to 1300 m. conservation measures. considered Vulnerable (VU) according to IUCN Red List criteria (IUCN, 2001). Additional ecological and and fruiting specimens were collected in May. 264 Novon with basal and cauline trifoliolate leaves, large leaflets with smooth margins, and the peduncles a: and flowers, a panicle with many flowers, two sepal with long pubescent hairs. whorls, and spurred petals. The new species clearly Cytological observations of Epimediun n both surfaces, the white inner sepals with 2E), indicate that E. yinjiangense if & Tian 2011 A New Species of Peperomia (Piperaceae) from the Cauca Valley, Colombia 2011. doi: 10.341' Volume 21, Number 2 2011 of 2 2011 Alpinia oui (Zingiberaceae), a New Species from Taiwan Yen-Hsueh Tseng National Chung Hsing University, Department of Forestry, 250 Kuokuang Road, Taichung, 402, Taiwan, tseng2005@nchu.eclu.tw Chih-Chiang Wang National Penghu University, Department of Tourism and Leisure, 300 Liuho Road, Makung City, Penghu County, 880, Taiwan, ccwang@npu.eclu.tw Abstract. Alpinia oui Y. H. Tseng & Chih C. resembles A. zenunbet (Pers.) B. L. Burtt & R. M. Sm. in the shape and size of the flowers, but differs in the cence of the capsules. According to the IUCN Red as Vulnerable (VU Dl), because its wild populations Key words: Alpinia, IUCN Red List, Taiwan, es 230 species that are mainly distributed in the and the Pacific Islands (Wu & Larsen, 2000); 51 2000) Thirteen taxa of Alpinia were previously recognized in the Flora of Taiwan (Moo, 1978); more edition of the Flora of Taiwan (Wang, 2000). Since been reported (Kuo et al., 2008; Liu & Wang, 2009; Tseng et al., 2010). During a recent revision of the Wang, 1998; Kuo et al., 2008; Liu & Wang, 2009; Liu et al., 2009; Tseng et al., 2010) and relevant 2000; Kress et al.. 2005; Chaveerach et al.. 2008) Alpi nia oui Y. H. Tseng & Chih C. Wang, sp. nov. TYPE: Taiwan. Taitung Co., Taimali township, Yaoshan. at forest margin, along semi-shaded Haec species Alpiniae zerumbet ( Pers.) B. L. Burtt & R to 1 cm; blades oblong to oblong-lanceolate, 30-60 X 7-15 cm, apex acuminate to obtuse, base cuneate. 2 cm, lateral lobes 2, basally 1/3 connate, oblong, ca. 3 X 1.2 cm; labellum ca. 3.5 X 2.5 cm, convolute, at style base, brown, ca. 2 mm; ovary green, densely the lower pedicels 0.5-1 cm: fruit capsular, red at margins at elevations of 500-600 m. Novon 21: 270-273. Published on 27 June 2011. doi: 10.3417/2009111 Volume 21, Number 2 2011 Tseng & Wang Alpinia oui (Zingiberaceae) 271 IIJCN Red List category. According to IUCN Red List criteria (IUCN, 2008), Alpinia oui is treated here as Vulnerable (VU Dl), with D1 indicating that the wild populations are small, with less than 1000 272 273 2011 Quercus delgadoana (Fagaceae), a New Species from the Sierra Madre Oriental, Mexico mce M. Kelly 3-5126, U.S.A. et al. 275 2011 276 Novon 2011 A New Species of Memecylon (Melastomataceae) from Taiwan Chih-Chiang Wang,1 Yen-Hsueh Tseng ,2 Yun-Tsao Chen,3 and Kun-Cheng Chang2* 'Department of Tourism and Leisure, National Penghu University, Penghu 880, Taiwan department of Forestry, National Chung Hsing University, 250 Kuo-Kuang Rd., 2011. doi: 10.3411 mil Volume 21, Number 2 2011 et al. Elat.ostema recurviramum (Urticaceae), a New Cave-dwelling Species from Guangxi, China Yi-Gang Wei State Key Laboratory of S; 1. The fertile axis a receptacle (Wang, 1980a, 1980b). Weddell (1869) & Moritzi) Miq. in CDBI, HITBC, II doi: 10.341) 2011. 282 Novon Wei & 2011 Two New Species of Aristolochia (Aristolochiaceae) from Hainan Island, China Hainan, IUCN Red List. al„ 1988; Huang et al„ 2003). 1. Aristolochia ledongensis Han Xu, Y. D. Li & H. J. Yang, Han Xu & H. Q. Chen JFL00972 CANT). Figure 1. nifoKaeC. F. Liang ex ; tlavo tubo basali ! 0.7-1 1.7-3. 9 cm, leathery. C 2-3 mm deep, apex 0.7-1 0.6 X tube 15-16 X 4.5-5 n: 2011. 286 Novon Xu et al. 201 1 ’ Aristolochia (Aristolochiaceae) from China Novon mi 2011 i. The paper was s People’s Republic of China (1 I Synonymy and Typifications in Groutiella tomentosa (Orthotrichaceae, Bryopsida) Yu Ning-Ning,1* Jia Yu ,2 2011. Yu et al. 2011 Volume 21, Number 2 2011 Yu et al. Volume 21, Number 2, pp. 161-294 of NOV ON was published on 27 June 2011. www. mbgpr ess . info CONTENTS Missouri Botanical Garden A Journal for Botanical Nomenclature VOLUME 21 NUMBER 3 2011 Volume 21 , Number 3 September 201 1 Novon, A Journal for Botanical Nomenclature from the Missouri Botanical Garden The mission of the Missouri Botanical Garden is to discover and share knowledge about plants and their environment, in order to preserve and enrich life. Novon publishes short articles whose primary purpose is the establishment of nomencla¬ ture in vascular plants and bryophytes. All articles are peer-reviewed by qualified, indepen¬ dent reviewers. Manuscripts must fully state and justify the reasons for proposing nova. These may include detailed comparisons with similar taxa, short keys to similar taxa, illustrations to similar taxa, and mechanical nomenclature reasons, among others. Manuscripts whose primary purpose is other than establishment of new nomenclature, which usually are longer manuscripts, cannot be accepted for review. These include reviews, revisions, monographs, or other papers that incidentally include nova. Manuscripts must follow the guidelines in the Checklist for Authors. The Checklist may be downloaded from the Garden’s web site, www.mbgpress.info, or authors may contact the man¬ aging editor to request a copy. Novon will not knowingly accept manuscripts that have been simultaneously submitted to other journals for consideration or previously published in some form elsewhere. Victoria C. Hollowed Editor Beth Parada Managing Editor Allison M. Brock Associate Editor Tammy Charron Associate Editor Cirri R. Moran Press Coordinator Ihsan A. Al-Shehbaz Consulting Editor Carmen Ulloa Ulloa Consulting Editor George Yatskievych Consulting Editor Kanchi N. Gandhi Nomenclature Consultant Nicholas J. Turland Nomenclature Consultant Roy E. Gereau Latin Editor Novon is included in the subscription price of the Annals of the Missouri Botanical Garden. Price for 2011: $180 per year U.S.A.; $190 Canada and Mexico; $215 all other countries. Four issues per Postal address Subscriptions Annats of the Missouri Botanical Garden, P.O. Box 1897, Law¬ rence, KS 66044-8897; Manuscript submissions: Novon, Missouri Botanical Garden, P.O. Box 299, St. Louis, MO 63166-0299. Web site: http://AAww.mbgpress.info © Missouri Botanical Garden 2011 © Novon is printed on paper that meets the re¬ quirements of ANSI/NISO Z39.48-1992. Novon (ISSN 1055-3177) is published quarterly by the Missouri Botanical Garden, 2345 Tower Grove Avenue, St. Louis, MO 63110. Periodicals postage paid at St. Louis, MO and additional mail¬ ing offices (USPS #006-777). POSTMASTER: Please send address changes to Annals of the Missouri Botanical Garden, P.O. Box 1897, Lawrence, KS 66044-8897. Information on the contents of Novon is present¬ ed in SciSearch®, Research Alert®, Current Con- tents®/Agriculture, Biology, and Environmental Sciences, APT Online, the CAB Abstract/Global Health database, and the ISI® database. The full-text of Novon is available online through BioOne™ (http://AVAVw.bioone.org). Volume 21 NO VON Number 3 VJK 2011 A New Species of Diploid Quillwort (. Isoetes , Isoetaceae, Lycophyta) from Lebanon Jay F. Bolin,1'2 Rebecca D. Bray,2 and Lytton John Musselman2* department of Botany. Smithsonian Institution, NMNH MRC 166, P.0. Box 37012, Washington D.C. 20013-7012, U.S.A. Current address: Department of Biology, Catawba College, 2300 W. Lines Si.. Sali-Lmx. \orlh Carolina 28144, U.S.A. bolinj@si.edu department of Biological Sciences, Old Dominion University, Norfolk, Virginia 23529-0266, U.S.A. rbray@odu.edu * Author for correspondence: lmusselm@odu.edu ABSTRUT A new species. Isoetes libanotua Mus¬ selman, Bolin & R. D. Bray (Isoetaceae, Lycophyta), 338 to 477 pm with remote, low tuberculate oma- species (Bolin et al., 2008). This paper describes an (K. Knio, pers. comm.). Recently, Musselman and Al- each in Turkey and Syria (Bolin et al., 2008). From sp. nov. TYPE: Lebanon. Akkar Distr.: 339 m, 34'37.163"N, 36'14.976"E, 13 Apr. 2009, M. S. Al-Zein & L. J. Musselman 2UU9-39c (holo- type, MO; isotype, BEI). Figure 1. Uth (10 to)20(to 50) sporophylls (leaves) (2-)4(— 6) doi: 10.3417/2010028 Novon 21: 295-298. Published on 9 September 2011. 296 Novon et al. 2011 Novon Isoetaceae) with terrestrial habitat in Calabria (Italy). New karvological and taxonomical data. FI. Medit. 11: 303-309. IlICN. 2001. IlICN Red List Categories and Criteria, Version 3.1. Prepared by the IUCN Species Survival Review of Chinese Spiraea (Rosaceae, Spiraeoideae) with Simple Inflorescences Roman Businskv Silva Tarouca Research Institute for Landscape and Ornamental Gardening (RILOG), 252 43 Pruhonice, Czech Republic, businsky@vukoz.cz stan and hitherto neglected in Chinese floras, is the ern Sichuan, S. muliensis T. T. Yu & L. T. Lu and S. Spiraea. some imperfectly known species, as well as the guished here. One species from each pair is found recent molecular analyses (Potter et al.. 2007) reveal Maximowicz (1879: 173). in his comprehensive rescences of true spiraeas as “flores haud vere paniculae compositae.” Schneider (1905: 449. 450. Euspiraea C. K. Schneid. (the genus Spiraea by type into one group with “Blutenstand doldig. ± doldentraubig, gestielt.” Rehder (1927: 333. 1940: 328) indicated the simple types similarly as doi: 10.3417/2009085 Novon 21: 299-316. Published on 9 September 2011. 300 Novon old classifications, e.g.. Maximowicz. 1879: Zabel. 1893; Schneider. 1905; Rehder. 1927. 1940) or a Nakai. 1916; modified by Pojarkova, 1939. and Yu & Kuan. 1963: and used in Chinese floras: Yu & Lu, basal leaf rosette: (2) shortly pedunculate (subses- sile). congested corymbs, with a basal leaf rosette; (3) Spiraea sect. Chamaedryon Ser. in a narrower concept (Yu & Kuan. 1963; Yu & Lu. 1974; Businsky & Businska. 2002). The term umbel-like and umbel (Rehder. 1927. 1940). Maxwell and Knees and corymb, and can be defined as an elongated carried out a series of research expeditions to China, where all taxa of the genus Spiraea that we found in nature were sampled (Businsky & Businska. 2002: 11). In addition to these cited specimens, most Spiraea collections at PE and KUN were also north-northwest of Lijiang. The latter name was today in different prefectures (Lijiang and Diqing, another. Yu and Kuan (1963) listed both taxa ser. Hypericifoliae Pojark. ex T. T. Yu Both taxa were & Lu. 1974; Lu & Crinan, 2003). and the same both publications (Yu & Lu. 1974: 61. t. 8; Wu & Raven. 2004: 34). Beyond the various leaf shapes were also accepted separately in the Index Florae given from between 2700 and 3600 m elevation, and Detailed study of the type material for both S. distinctly angled, dark brown shoots and branchlets. Spiraea calcicola W. W. Sm., Notes Roy. Bot. Card. Edinburgh 8: 131. 1913. TYPE: China. stone cliffs on the eastern flank of the Lichiang Range [Yulongxue Shan]. 27°20'N [27°10'N], 11.000-12.000 ft.. June 1910. G. Forrest 5730 (holotype. E not seen; isotype. PE). Figure 1. P 13, Tig. 1. 1933, syn. nov. TYPE: China. Yunnan: montis Lamatso. Substr. Calceo, ca. 3200 m,” 12 July 1915, Handel-Mazzetti 7608 (holotype, W). Volume 21, Number 3 30403 (G), B-D from isotypes. 2011 but often obtuse at very base, (15-)20-30(-35) X 7- 17 pairs in upper third of narrower leaves on short fertile branchlets or in lower part of sterile long shoots; pinnate with usually 3 pairs of lateral veins leading to 5, with ca. 10 to 15 r one area in the c Yushan (Mt. Jade. 3952 m). S. tatakaemu is near the border of Chiayi and f Yu and Lu (1974), S. tarokoemis is placed within the 306 Novon 2011 & Su, 1977; Ohashi & Hsieh, 1993). The il in the new Flora of Taiwan treatment (( Hsieh. 1993: 156. pi. 71) is ascribe a (Hayata ex Nakai) H. L. Li. There is 1.2-2X 1.7-2. 4 mm. [Taroko Natl. Park], Kwarenko, B. Hayata & S. Sasaki 68 (TAIF 12535, 12536 [s.n.], 12537); [Taroko Natl. Park] Yenhai logging trail, Shih-Wen Chung 8006 (TAIF), Pi- 111111^= Flora of China Pedicels Volume 21, Number 3 2011 Businsky Review of Chinese Spiraea (Rosaceae) densely whitish tome deep green adaxially; shallowly crenate to central to northeastern mostly 10-20 central to southeaster! entirely glabrous), and by the hypanthium and sepals Schneider, 1905; Rehder, 1927, 1940; Bean, 1981), or in Chinese floras (Yu & Lu, 1974; Lu & Crinan, Pojarkova. 1939; Shul’gina. 1954; Svjazeva. 1980) 1972) and India (Purohit & Panigrahi, 1991). The Mongol, and as presented in Chinese floras (Yu & Lu, 1974; Fu & Hong, 2003; Lu & Crinan, 2003). (Schneider. 1905; Rehder. 1913. 1927. 1940; Bean. 1981; Maxwell & Knees, 1995) for the relevant northwestern Himalayas to north-central China is S. 310 Novon 2011 Jiulong \ian, bottom oi uppei Taka He t allot, 1100 n 29°01 'N, 101°49'E, 5 July 2001, R. Bminskf 51405 (t G. Gmelin s.n. (t Notes. The name Spiraea conjusa Regel & Korn, published in 1857, has been used in the older Acta Hort. Petrop. ( i-lc- Mo-eou 10: 501. 1813. TYPE: [Russia (Chita Province) & China (NE Nei Mongol) border.] “Dahuria. in lapidosis ad fluvium Svjazeva, 1980) and in Chinese floras (Yu & Lu, 1974; Lu & Crinan, 2003; Fu & Hong, 2003); it differs from typical S. media only by leaf blades laxly of China key. Alto W. W. Sm. 2011 314 2011 A New Species and One New Name in Castanopsis (Fagaceae) from Hainan, China Li Chen , Zi-Gang Zhang, Ying Hu, Xin-Wei Li,* and Jian-Qiang Li* Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, Hubei 430074, People's Republic of China * Author for correspondence: lijq@rose.whiob.ac.cn M. Chen & B. P. Yu) J. Q. Li & Li Chen (Fagaceae), sandy beaches. Castanopsis gl.abrifolia is closely on its glabrous branches and petioles, the smaller nov. TYPE: China. Hainan: Wenchang, Chang- 14 Jan. 1978, Anonymous 9254 (holotype, IBSC 0034559). Figure 1. Chen & B. P. Yu, J. !.' China Agile. Univ. 12: 93. 1991, syn. nov. TYPE: China. Hainan: Wenchang, 11 Dec. 1987, B. P. Yu 103158 (holotype, CANT). (D. Don) Spach (Fagaceae). Camus (1929) recorded by Barnett (1944), who recognized 119 species in 11 groups. Govaerts and Frodin’s (1998) World Checklist China. Most recently, Huang and Chang (1998) and Huang et al. (1999) revised Chinese Castanopsis and accepted 63 species and 58 species in the Flora several decades (Fu & Huang. 1989: Huang & Chang, 1990, 1996; Chen & Yu, 1991; Fu & Feng, 1992; Fu, 2001; Chen et al., 2009, 2010). During red-brown after cortices dehisce, lenticels slightly 8.5(— 10) X 1.1 — 3( — 4.5) cm, leathery and concolor- secondary veins in 6 to 9 pairs on each side of 3.5-7 X 0.1-0.15 cm; cupule prolate ellipsoid, 1.1- splitting into 2 to 3 segments when mature, cupule transversely united across 3 or 4 rings, the external pubescence and scales. Fruit as 1 nut per cupule, apex, ca. 1.2 X 0.6-0. 8 cm, scar basal, slightly doi: 10.3417/2009103 Novon 21: 317-321. Published on 9 September 2011. 318 Novon Garden, CAS) from Anonymous 9254 (IBSC). IUCN Red List, category. Castanopsis glabrifolia is known from four populations from Wenchang Endangered (CR) according to IUCN Red List Categories and Criteria (IUCN, 2001). Etymology. The specific epithet glabrifolia is taken from the Latin and refers to the glabrous leaves of the new species. Chen et al. 2011 Chen et al. 2011 Inga pitmanii (Fabaceae), a New Species from Madre de Dios, Peru doi: 10.341' 2011 pedicel 0.5-1 mm. puberulent. Flowers with a 2011 325 New Combinations Within the Briza Complex (Poaceae, Pooideae, Poeae) Liliana Essi Universidade Federal de Santa Maria, Centro de Educatjao Superior Norte do Rio Grande do Sul, Departamento de Zootecnia e Ciencias Biologicas, Av. Independence 3751, Bairro Vista Alegre, 98300-000 Palmeira das Missoes, Rio Grande do Sul, Brazil, lili.essi@gmail.com Hilda Maria Longhi Wagner and Tatiana Teixeira de Souza Chies Universidade Federal do Rio Grande do Sul, Instituto de Biociencias, Departamento de Botanica, Av. Bento Gontjalves 9500, Bairro Agronomia, 91501-970 Porto Alegre, Rio Grande do Sul, Brazil. hmlw@plugin.com.br, tatiana.chies@ufrgs.br Abstract. Briza L. (Poaceae, Pooideae, Poeae) is a 1987), or sensu stricto, including only four Eurasian (1975) and other authors. The American species. Wagner & Souza-Chies, C. brasili-ense (Nees ex brizoides (Lam.) Essi, Longhi-Wagner & Souza- Wagner & Souza-Chies, C. itatiaiae (Ekman) Essi, (Trin.) Essi, Longhi-Wagner & Souza-Chies, C. monandrum (Hack.) Essi, Longhi-Wagner & Souza- (Nees) Essi, Longhi-Wagner & Souza-Chies. The The genus Briza L. (Linnaeus, 1 75.'!: Poaceae, added later, in 1808. Several new South American (Nicora & Rugolo de Agrasar. 1981). Further 1920; Rosengurtt et al., 1968, 1970; Longhi-Wagner. Matthei (1975) published a revision of Briza, and the Eurasian species, and distributed the American authors (Smith et al., 1981; Soreng et al., 2003). Nicora and Rugolo de Agrasar (1981) described (Ekman) Parodi ex Nicora & Rugolo and M. poomorpha (J. Presl) Parodi ex Nicora & Rugolo [as Agrasar (1981) accepted Briza with a different (1975), including not only the Eurasian species, as Novon 21: 326-330. Published on 9 September 2011. doi: 10.3417/2010026 2011 2011 Nomenclatural Changes for Zornia (Leguminosae, Papilionoideae, Dalbergieae) in Brazil Ana Paula Fortuna-Perez Departamento de Biodiversidade, Evolutjao e Meio Ambiente, Universidade Federal de Ouro Preto, Campus Morro do Cruzeiro-ICEB, Ouro Preto, Minas Gerais, CEP 35.400-000, Brazil Author for correspondence: paulaforperez@yahoo.com.br Ana Maria Goulart de Azevedo Tozzi. Departamento de Biologia Vegetal, Universidade Estadual de Campinas, P.0. Box 6109, Campinas, Sao Paulo, CEP 13.083-970, Brazil Chodat & Hassl.. /,. diphylla var. elatior Benth.. Fort. -Perez & A. M. G. Azevedo is proposed, based Mohlenbr. Eighteen new synonymizations are also Key words: Brazil, Dalbergieae, Leguminosae, Neotropics. Papilionoideae, Zornia. brocU 1961; Rudd, 1981; Klitgaard & Lavin, 2005). informal Adesmia DC. clade of the tribe Dalbergieae nuclear ribosomal 5.8S and flanking internal tran- flower, the 2- or 4-foliolate leaves, and the stems that (1791) based on Z. bracteata J. F. Gmel/ After Desv., describing M. tetraphyllus (L.) Desv. De accepted this distinction. However. Vogel (1830) was later followed by Bentham (1859), Moricand (1844), and others. More than 30 species of Zornia eight species were accepted, divided between the two various species with Z. diphylla (L.) Pers. and by Mohlenbrock (1961), who recognized two infra- Fortuna-Perez (2009). more than 3000 accessions were analyzed through visits to European and ing AHUC, B, BHCB, BM, BR, CEN, CEPEC, CGMS, CTES, ESA, F, FHO, G, GH, H, HRCB, HST, HUEFS, IAC, IAN, IBGE, ICN, INPA, INTA, IPA, K, LIL, M, MBM, MEXU, MG, MICH, MO, MVM, NY, OXF, P, PACA, PAMG, PEUFR, RB, S, SI, SP, SPF, UB, UEC, US, W, Y, and Z. As part of this revision. 36 species were recognized, eight doi: 10.3417/2010040 Novon 21: 331-337. Published on 9 September 2011. !Ii if Volume 21 , Number 3 Fortuna-Perez & Azevedo Tozzi 335 201 1 Nomenclatural Changes for Zornia (Leguminosae) Zornia ovata Vogel, Linnaea 12: 58. 1838, syn. nov. TYPE: Brazil. Minas Gerais: Datas, 28 Oct. 2006, A. P. Fortuna-Perez et al, 125 Ineotvpe, designated here, UEC). Zornia diphylla (L.) Pers. var. pubescent (Kunth) Benth., FI. Bras. (Martins) 15(1): 82. 1859, syn. nov. Basionym: 515. 1823. TYPE: Colombia. Fusagasuga, s.d., A. J. iholotvpe, B, II - \\ 13775). 1838, syn. nov. TYPE: Brazil. Id'., F. Sellout s.n. (holotype, K). Notes. When Smith (1818: 205) described Zor- collection: “ Hedysarum diphyllum (5: Willd. Sp. PI. v. 3. 1178. 8; Lamarck v. 6.404. //. diphyllum: Swartz Obi. 285. H. n. Browne Jam. 301. excluding the Redhead (1963). Smith (1818) possessed the Lin- naean Herbarium, which contained Patrick Browne’s 921.23 that is deposited in the LINN Herbarium was Rees’s Cyclop. [[5] Willd.” and annotated by his treatment, Mohlenbrock (1961) mistakenly estab¬ lished a neotype ( Harris 12070 [US], from Jamaica), Dandy and Milne-Redhead (1963) also cited LINN 921.23 in the LINN Herbarium is being designated ovata, which were treated by Mohlenbrock (1961) as described by Vellozo (1825). When Mohlenbrock and Z. pubescens as synonyms of Z. latifolia. his When Vogel (1838: 58) described Zomia ovata, he perforata (synonymous with Z. glabra). The original widely from Z. sencea. Because none of Sellow’s material that correspond¬ ed to Zomia ovata has been located, the type (Brazil. for this Vellozo name, and indeed for many names in Florae Fluminensis. It can be assumed that the plate author’s intent (McNeill et al., 2006: Art. 9.10). The plate was likely based on a collection that was lost When Chodat and Hassler (1904) described and Hassler (1904) described Z. diphylla f. diversi- 11. Zornia sericea Moric., PI. Nouv. Amer. 126-127. 1844. TYPE: Brazil. Bahia: Jacobina, s.d., J. S. duplicates. BM. K. Ny! P. W). ? of Z. ovata Vogel (Vogel, 1838). However, in this tion of Z. serieea, which is reestablished here. Zomia bract coles (vs. lanceolate), the length of the auricles at the bracteole (0—3 mm vs. 6-10 mm), three to four A New Species of Gaultheria (Ericaceae) from Mount Kinabalu, Borneo, Malaysia Valdosta State University, 1500 N. Patterson 31698-0015, U.S.A. cmbush@vaklosta.edu Volume 21, Number 3 2011 Fritsch & Bush Gaultheria (Ericaceae) from Borneo 339 340 2011 keeled, 1.5-2 X 1.2-1. 5 mm. otherwise similar to bracts. Calyx 3-4 mm; lobes 5, white tipped with dark pink, ovate. 2-2.5 X 1.7-2 mm, outside . 4-4.5 X 3-3.5 i 0.7-1. 5 mm; stamens 10; filaments 1.2-1.4 mm. 0.5-0. 6 1.5-2. 5 a prostrate shrub habit to 0.6 m tall (vs. shrub to 2.4 m), leaf blades 0.8-1.5(— 2) X cm (vs. 1.8-2 .4 X 0.6-1 cm), only one 0.3-0. 7 G. 0.3 bract eoles not keeled' and with the margins not o lobes 1-1.8 X fr.), W. Hoya jianfenglingensis (Apocynaceae), a New Species from Hainan, China Shao-yun He College of Hoilieulluie, Smlli ( Miiiia Agricultural University, Guangzhou 510642, People's Republic of China, syhe2001@163.com Ping-tao Li * and Jia-yi Lin College of Forestry, South China Agricultural University, Guangzhou 510642, People's Republic of China Guoyang Lin Pennsylvania State University, 221 Shields Building, University Park, Pennsylvania 16802, U.S.A. gyll@psu.edu Hui-lan Zeng College of Hoilieulluie, ^-oii I h China Agricultural University, Guangzhou 510642, People's Republic of China *Author for correspondence: Li85280485@126.com Abstract. Hoya jianfenglingensis Shao Y. He & P. with the related species. //. fimgii Merr. and //. camosa (L. f.) R. Br. Hoya jianfenglingensis differs in six to 10 pairs, the evident glands at the corona base, the genus on the basis of the acute outer angle of the Hoya, sect. Hoya, IUCN Red List. distributed in tropical and subtropical areas from eastern Asia throughout the Pacific archipelago and forms were reported from China (Tsiang & Li, 1977), but Li et al. (1995) later considered 32 species and a Shao Y. He & P. T. Li a recent addition to Hoya of China (He et al., 2009). Most species occur in are 22 species with a white corolla in China. Recently, Forster and Liddle (1996) estimated the genus at over 300 species, distributed in Malaysia. 29 species noted for India. Although Rintz (1978) agreed with Ridley (1923) on a species total of 25 Hoya in Malaysia, they significantly differed in five Pakistan (Nasir & Ali. 1972). During two expeditions to Jianfengling, Hainan, in the autumn of 2006 and summer of 2008. three HITBC, IBSC, and KUN, and study of the pertinent literature (Hooker, 1885; Costantin, 1912; Merrill, 1932; Nasir et al., 1972; Tsiang & Li, 1974, 1977; Rintz, 1978; Li, 1994; Gilbert et al., 1995; Li et al., 1995; Foster & Liddle, 1996; Lu et al., 1998; Jagtap doi: 10.3417/2009061 Novon 21: 343-346. Published on 9 September 2011. 344 Novon Y. Li & J. Y. Lin 0607281 (holotype, CANT, "Ique inis ad ‘ ba4m CANT photo at MO). Figure 1. glanduliieria differ!. He et al. Hoya (Ap 2011 Senecio ser. Chilenses , a New Name for Senecio ser. Suffruticosi (Asteraceae, Senecioneae) from Southern South America Mariana Gabriela Lopez and Arturo Federico Wulff Laboratorio de Citogenetica y Evolucion, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Lit. Giiiraldes 2620, 1428 Buenos Aires, Argentina, and Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET). magalo@ege.fcen.uba.ar; artulf@ege.fcen.uba.ar Cecilia Carmen Xifreda Laboratorio de Etnobotanica y Botanica Aplicada, Facultad de Ciencias Naturales, and Museo, Universidad Nacional de La Plata, calle 64 N° 3, 1900 La Plata, Argentina, cxifreda @fcnym . unlp . edu . ar Abstract. The name Senecio L. ser. Chilenses DC. ex M. 0. Lopez, A. F. Wulff & Xifreda is here epithet Suffruticosi was published at different cioneae) for two different taxa from South and North Resumen. El nombre Senecio L. ser. Chilenses DC. ex M. G. Lopez, A. F. Wulff & Xifreda se valida aquf rangos subgenericos diferentes dentro Senecio (Aster- Our studies on the Asteraceae of Andean high tural illegitimacy of the name Senecio L. ser. Suffruticosi Cabrera (Cabrera. 1985). It is a later Code of Botanical Nomenclature (ICBN: McNeill et al., 2006: Art. 53.4). Therefore, we herein validate Lopez, A. F. Wulff & Xifreda in replacement. 1970). Greenmail (1915) also recognized his section 1970). Independently, Cabrera (1949) published the later reduced the taxon to the rank of series, publishing the new epithet Suffruticosi Cabrera, agreement with ICBN Art. 21.2, stating that the name agreeing in gender with the generic name” (McNeill et al.. 2006: 42). Argument might be made that the epithet Suffrutecius (Cabrera, 1949) is an orthograph- Art. 53.3 (McNeill et al.. 2006). an orthographical examples differing by only one or two letters, and the (1985: 194) published his "series. Suffruticosi as a Cabrera, Lilloa 15: 101. 1049. •Suffrutecius.’” in conformity with ICBN Rec. 50F (McNeill et al., 2006). In addition, later in Cabrera et al. (1999), he again cites “sect. Suffrutecius Cabrera (1949),” Suffruticosi (Cabrera. 1985). doi: 10.3417/2008127 Novon 21: 347-348. Published on 9 September 2011. Musella lasiocarpa var. rubribracteata (Musaceae), a New Variety from Sichuan, China Ma Hong,1 Pan Qingjie ,2 Wang Lan,3 Li Zhenghong,1* Wan Youming,1 and Liu Xiuxian1 'Research Institute of Resource Insects, Chinese Academy of Forestry, Kunming 650224, China 2Shaohai Development Management Office of Jiaozhou City, Shandong 266300, China 3Yunnan Reascend Tobacco Technology (Group) Co., Ltd., Kunming 650106, China *Author for correspondence: hortscience@163.com Abstract. Taxonomic study of the diversity of Musella lasiocarpa (Franch.) C. Y. Wu ex H. W. Li Musella lasiocarpa (Franch.) C. Y. Wu ex H. W. Li represents the monotypic genus Musella (Franch.) C. Y. Wu & H. W. Li in the family Musaceae (Li, 1978, 1979; Wu & Kress, 2000). The taxon is endemic to recognizable one within the Musaceae, distinguished 5 cm. This attractive plant has been horticulturally was eventually recognized (Cheesman, 1947; Sim¬ on ITS and trnL-F sequence data supports the status Musella should be a distinct genus (Xue et al., 2007). known to exist on cliffs within the watershed of the upper Yangtze River and its tributaries, in northern locally common in this region of southwestern China Long et al.. 2008). committed to investigate the diversity of wild 1. Musella lasiocarpa (Franch.) C. Y. Wu ex H. W. Li, Acta Phytotax. Sin. 16(3): 56-57. 1978. (Morot) 3(20): 330-331. fig. 1. 1889. Ensete 2(2): 102. 1947. TYPE: Fig. 1 in Franchet. Vare. 2008: 81). doi: 10.3417/2010125 Novon 21: 349-353. Published on 9 September 2011. 350 Novon 18)/phot(fby H. Ma and Q. J. Pan. 2011 Ma et al. larly ridged tips ‘(Fig. 1J. K). Chromosome number: China. The 352 Novon Ma et al. 2011 A New Combination in the Fern Genus Osmundastrum (Osmundaceae) William A. McAvoy Delaware Natural Heritage and Endangered Species Program, Delaware Department of Natural Resources and Environmental Control, Division of Fish and Wildlife, 4876 Hay Point Landing Rd., Smyrna, Delaware 19977, U.S.A. william.mcavoy@state.de.us glandidosum (Waters) McAvoy. Key words: IUCN Red List, North America, (2008) and Jud et al. (2008) confirmed the work of earlier studies (Tagawa, 1941: Hewitson. 1962; Bobrov, 1967; Miller, 1967, 1971; Yatabe et al., 1999, 2005) that the genus Osmunda L. is para- Osmundastrum C. Presl at the level of genus and as its only extant species. In 1902, Campbell Easter Waters (1872-1955) guished by its densely glandular pubescent foliage as distinctiveness of the taxon. The varietal name is didosa Waters. Fern Bull. 10: 21-22. 1902. TYPE: U.S.A. Maryland: Glen Burnie, low wet woods with the typical, 17 July 1901, C. E. 079887). were stated to be at the “National Museum. Gray Herbarium” (Waters, 1902: 22). and syntypes were 1950) recognized this variety, as did Reed (1953) ami invalidly as a form (Broun, 1938;^Weakley, 2010), synonymy (Whetstone & Atkinson, 1993), or do not consider it (Gleason & Cronquist, 1991). Wagner (1991: 20) was intrigued with this taxon and stated: Femald (1942: 353) first encountered this species in Virginia, he noted that: “whenever we took hold of Waters (1902: 21) recognized that both the glandular-pubescent variety and the typical variety distinctiveness: “Both the variety and the typical both the glandular and nonglandular varieties covered with stiff, glandular hairs ( < 1 mm long) that Novon 21: 354-356. Published on 9 September 2011. doi: 10.3417/2010045 A New 2011 The Identity of Synandropus and a New Combination in Neotropical Menispermaceae Rosa del C. Ortiz Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166, U.S.A. rosa.ortiz-gentry@mobot.org within section Somphoxylon (Eichler) Bameby. The Ortiz is made: 0. uva-alba Bameby and 0. arifolia Resumen. Se lectotipifica Synandropus membrana- ceus A. C. Sm. (Menispermaceae), conocida ante- rionnente solo de la coleccion tipo y se reduce Miers, en la section Somphoxylon (Eichler) Bameby. (A. C. Sm.) R. Ortiz: 0. uva-alba Bameby y 0. arifolia Bameby so >, Odon- with three orders of branching and a 3-merous sepals, six fleshy petals, and a 3-merous synandrium (Bameby, 1970). Mem. ^New York Bot. Card. 20(2): 110. 1970. Basionym: Somphoxylon Eichler, Flora 47: 396. 1864. TYPE: Somphoxylon wullschlaegelii Eich- Bameby]. Synandropus A. C. Sm., Bull. Tonev Bot. Club 58: 93. 1931, syn. nov. TYPE: Synandropus membranaceus A. ceus A. C. Sm.. Bull. Torrey Bot. Club 58: 93. 1931. TYPE: Brazil. Para: Breves, Amazon estuary, swampy jungle, 23 Oct. 1929 (c? inflor.), E. P. Killip & A. C. Smith 30217 (lectotype, designated here, NY 00320662 [c? 0010973 [c? inflor.. leafless branch at left]). Figure 1. 20(2): 120. 1970, syn. nov. TYPE: Pem. Loreto: Ayuavtia, in forest, 250 m, 20 Mav 1959 (fr.), F. Woytkowski 5374 (holotype, F 1532620; isotype, M0 1744787). 22(4): 142. 1972, syn. nov. TYPE: Pem. San Martin: in forest thicket, Saposoa, 14 Oct. 1959 (fr.), F. 3459413 [fruiting branch]!' duplicates, M0 2143393 lectotype (Fig. 1A) correctly belongs to the Menis- are clearly allied to the Euphorbiaceae. The doi: 10.3417/2010092 Novon 21: 357-361. Published on 9 September 2011. S-uJU-^ 358 Novon Volume 21, Number 3 2011 Ortiz Neotropical Menispermaceae 359 361 2011 A New Species of Hechtia (Bromeliaceae) from Chihuahua, Mexico Ivon M. Ramirez Morillo Herbarium CICY, Unidad de Recursos Naturales, Centro de Investigacion Cientffica de Yucatan, A.C., Calle 43 #130, Colonia Chuburna de Hidalgo, Yucatan 97200, Mexico. ramirez@cicy.mx Adolfo Espejo-Sema and Ana Rosa Lopez-Ferrari Departamento de Biologia, Division de Ciencias Biologicas y de la Salud, Universidad Autonoma Metropolitana-Iztapalapa, Apartado Postal 55-535, 09340 Mexico D.F., Mexico. aes@xanum.unam.nix Abstract. Hechtia edulis I. Ramirez. Espejo & Lopez-Ferr. (Bromeliaceae) is described and illus- floral bracts scarious. and pistillate flowers with Parque Nacional Barranca del Cobre. Chihuahua, Mexico (ca. 94%; Espejo-Sema et al., 2004). All the ecious species from Baja California. Plants of Hech tia calcareous, volcanic, or chalky soils. stenopetala Klotzsch and //. schottii Baker, show that are odorless. In only a few instances are the and pistillate inflorescences (i.e., //. rosea E. Morren Hechtia has traditionally been considered within of the genus (Duvall et al., 1993; Homes et al., 2000; Reinert et al., 2003; Givnish et al., 2004), the within the family are uncertain (Givnish et al., 2007). str., as the sister group of Tillandsioideae (Homes et al., 2000; Crayn et al.. 2004). More recently, the analyses to date, and there is far from a satisfactory ongoing, and as a result extensive fieldwork has Novon 21: 362-367. Published on 9 September 2011. doi: 10.3417/2009130 etal. 2011 al„ 2007), //. pmiosa Espejo & Lopez-Ferr. and //. zamudioi Espejo, Lopez-Ferr. & I. Ramirez (Espejo- Serna et al„ 2008), //. lepidophylla I. Ramirez s ca. 1.2 X 2 cm, c 0.5-0. 7 364 Novon Volume 21, Number 3 2011 Ramirez Morillo et al. Hechtia (Bromeliaceae) from Mexico 365 366 Novon (diam.), almost sessile, peduncle conic, thick, ca. 1- shape, but variable among flowers in the same the central nerve; sepals free, widely triangular, fleshy, green basally, apically light brown and dry in ovate. 4-4.5 X 2.8-3 mm. apex acute, concave, green, fleshy, 3- to 5-nerved, sometimes 1 or 2 lateral (diam.), green, the placentation central, ovules orthotropous. white. 0.5-0. 7 mm; stigmatic lobes Barranca de Batopilas and Barranca de Candamena, Chihuahua State in the Sierra Madre Occidental. 67 municipalities in the state of Chihuahua (Espejo- Individuals of this species grow as lithophytes on oak-pine forest, formin g small colonies of one to four IIJCN Red List category. Hechtia edulis is only Data Deficient (DD), according to IUCN Red List criteria (IUCN, 2008). 8855) during a stay at the Missouri Botanical Garden staminate plants. A year later, a collecting trip was Barranca de Batopilas during September 2005. We few rosettes and were eventually able to flower that the leaves of Hechtia edulis are eaten by the Tarahumara people (fide Bye 8855). and flowers may greatly differ. As for many taxa in the genus, the affinities of the new species are of the rosettes place it in a group of species lapostollei L. B. Sm. and //. lyman-smithii Burt-Utley population (Burt-Utley & Utley, 1987). etal. 2011 A New Awned Species of Paspalum (Poaceae, Panicoideae, Paniceae) from Brazil Desiree M. Ramos V Jose F. M. Vails V2 Regina Celia de Oliveira V* and Dalva 370 Novon 371 2011 SelrS ms si!::: i clade by Denham et al. (2002) as clades A ; 372 A New Combination in Spiraea (Rosaceae) from Ulleung Island, Korea Hyunchur Shin Department of Biology, Soonchunhyang University, Asan 336-745, Republic of Korea. shinhy@sch.ac.kr Young-Dong Kim Department of Life Science, Hallym University, Chuncheon 200-702, Republic of Korea. yclkim@hallym.ac.kr Sang-Hun Oh L. H. Bailey TTorlomim. Department of Plant Biology, Cornell University, Ithaca, New York 14853, U.S.A. Author for correspondence: so253@cornell.edu Abstract. A rare endemic species on Ulleung morphology and DNA sequence data. The new raea, Ulleung Island. about six species in Rosaceae (Oh & Potter, 2005). simple leaves, fused carpels at least at the base. and one in eastern North America: the eastern North cultivated as an ornamental. In East Asia. P. China, and the Far Eastern Federal District of Physocarpus insularis (Nakai) Nakai has been volcanic island is located in the East Sea of Korea 1998). ulmifolia (Scop.) Maxim, by Kim et al. (2000), a distinct species in Physocarpus (Chung, 1957; Lee, DNA sequences (Oh et al., 2010). Spiraea L. (Oh et al., 2010). Comparative morphology characters of Spiraea. Leaves are unlobed and do not Phylogenetic analyses of chloroplast DNA (cpDNA) regions ( rbcL , matK, ndhF, and tmL-tmF) with species-level samples of Spiraea suggested that P. Spiraea chamaedryfolia by the size and shape of the leaf blade, the type of leaf base, and the number of insularis are significantly larger than those of S. broadly ovate and subcordate. truncate, or occasion- doi: 10.3417/2009126 Novon 21: 373-374. Published on 9 September 2011. 374 Manglietia crassifolia (Magnoliaceae), a New Species from Vietnam 376 Novon Volume 21, Number 3 2011 Vu et al. 377 378 Vu et al. 2011 Davallia napoensis, a New Species of Davalliaceae from Guangxi, China Wang Fa-Guo, Chen Hong- Feng, and Xing Fu-Wu Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, 510650 Guangzhou, Guangdong, People's Republic of China. Author for correspondence: xinfw@scib.ac.cn treatment of Davafliaceae for the Flora of China, it sis F. G. Wang & F. W. Xing is herein described and lamina 20-24 cm long, and the widely crateriform Red List. The genus Davallia Sm. (Davalliaceae), with about southeast to Polynesia (Nooteboom, 1996; Wu, 1999; Von Konrat et al., 1999). Kato (1985) provided a very detailed history of the generic delimitation until that time. The family Davalliaceae was historically classified into eight genera by Copeland (1908, 1927). Later, Kato (1985) recognized seven genera, cation, the generitype for Humata Cav. ill. ophio- Humata [= Davallia sect. Humata (Cav.) C. Presl], are included in Davallia sect. Wibelia. In 1989, Kato 1996). More recently. Kato and Tsutsumi (2008) Davallia and section Trogostolon (Copel.) M. Kato & China (Ching et al., 1959; Wu, 1999; Wang & Xing, 2008 i. Davalliaceae for the Flora of China, Volume 3, Davallia napoensis F. G. Wang & F. W. Xing, sp. nov. TYPE: China. Guangxi Prov.: Napo Co., Baidou, Nongbulin. near peak of evergreen broad-leaved forests, ca. 1200 m, 20 May 1989, South China Expedition 841 (holotype, IBSC). Figure 1. obtusis marginibus glabris, stipite 20-24 cm longo, lamina triangular-ovate. 25-28 X 18-22 cm. base nearly upward, spaced closely together, alternate or basal 1 cm long, bipinnate: pinnules in 10 to 12 pairs, leaflet larger. 0.9-1.6 X 0.5-1 cm. ovate-triangular. Novon 21: 380-384. Published on 9 September 2011. doi: 10.3417/2009093 Volume 21, Number 3 2011 Wang et al. Davallia napoensis from China 381 Volume 21, Number 3 2011 383 Petrocosmea isis, a New Species of Gesneriaceae from ; Areas in Guangxi, China Wei-bin Xu , Bo Pan , and Yan Liu* Wei & Wen, 2009), we c 1, 1.5-3. 5 X 1.5-2. 5 cm. 21: 2011. 386 Novon Xu et al. 2011 Fallopia multiflora var. angulata , a New Combination in the Polygonaceae from China Yan Hanjing,1'2 Fang Zhijian,2 Zhang Hongyi,2 and Yu Shixiao1* 'School of Life Sciences/State Key Laboratory of Biocontrol, Sun Yat-sen University, Guangzhou 510275, People's Republic of China 2School of Traditional Chinese Medicine, Guangdong Pharmaceutical University, Guangzhou 510006, People's Republic of China *Author for correspondence: lssysx@mail.sysu.edu.cn (Polygonaceae) reveals that a new combination in Fallopia Adans. is necessary, and the name F. Liu) H. J. Yan. Z. J. Fang & Shi Xiao Yu is proposed. In The Polygoneae of Eastern Asia, Steward (1930) divided the family into three genera (Antenoron Raf., Polygonum L., and Koenigia L.), and P. multiflorum Meisn. Liu (1959) later moved P. multiflorum along dent genus (Haraldson, 1978). and P. multiflorum (1998; Li et al.. 2003) in the Flora Reipublicae Haraldson is a well-known herbal medicine in China, (1991) described P. multiflorum var. angulatum S. Y. typical variety in its square branchlets with longitu- along the vein and abaxially. turning green at maturity: the larger flower (3-3.5 mm diarn. at and the elliptic tepals that are unequal in size, with China treatment of the Polygonaceae (Li, 1998). We examined the bioactive components of Polyg- (Thunb.) Haraldson var. multiflora ] collected from Tianyang County, Guangxi (Table 1). The bioactive ably from those of the typical variety (Fu et al., 2006; Bot. Upsal. 22(2): 77. 1978. Basionym: "jPo/y- 379. 1784. TYPE: China. Yunnan: Duel 'owe 485 (holotype, BS [fide Steward, 1930] not seen). lb. Fallopia multiflora var. angulata (S. Y. Liu) H. Basionym: Polygonum multiflorum Thunb. var. angulatum S. Y. Liu, Acta Bot. Yunnan. 13(4): 390. 1991. TYPE: China. Guangxi: Lingyun, 23 Oct. 1989, S. Y. Liu L89015 (holotype, KUN). Yonekitra & H. Ohashi. J." Jap. Bot. 72(3): 158. 1997. Basionym: Pleuropterus ciliinervis Nakai, Repert. Spec. Nov. Regni Veg. 13(363 — 367): 267-268. 1914. Fallopia multiflora var. ciliinewe (Nakai) A. J. Li, FI. Reipubl. Popularis Novon 21: 388-391. Published on 9 September 2011. doi: 10.3417/2009032 Yan et al. 389 2011 390 Novon g. — B. Fallopia midtiflora var. angulata (S. Y. Liu) H. J. Yan, Z. J. Fang & Shi Xiao Yu, ,S. Y. Liu Y05TY from Tiangyang, Guangxi. C-F. Light micrographs of root tuber transverse sections. C, D. and the typical variety of F. multiflora. The former Billlffllilli Yan et al. 391 2011 2 of NOVON was p www. mbgpr ess . info CONTENTS 317 331 338 343 347 349 354 357 362 368 373 375 385 Missouri Botanical Garden A Journal for Botanical Nomenclature VOLUME 21 NUMBER 4 2011 Volume 21 , Number 4 December 201 1 Novon, A Journal for Botanical Nomenclature from the Missouri Botanical Garden The mission of the Missouri Botanical Garden is to discover and share knowledge about plants and their environment, in order to preserve and enrich life. Novon publishes short articles whose primary purpose is the establishment of nomencla¬ ture in vascular plants and bryophytes. All articles are peer-reviewed by qualified, indepen¬ dent reviewers. Manuscripts must fully state and justify the reasons for proposing nova. These may include detailed comparisons with similar taxa, short keys to similar taxa, illustrations to similar taxa, and mechanical nomenclature reasons, among others. 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Information on the contents of Novon is present¬ ed in SciSearch®, Research Alert®, Current Con¬ tents®^ It B Ip IE n ntal Sciences, APT Online, the CAB Abstract/Global Health database, and the ISI® database. The full-text of Novon is available online through BioOne™ (http://www.bioone.org). Volume 21 NO VON Number 4 VJK 2011 New Species and Notes on Begonia (Begoniaceae) from Middle America, I Kathleen Burt-Utley and John F. Utley Department of Biological Sciences, University of New Orleans, New Orleans, Louisiana 70148, U.S.A. Current address: Department of Cell Biology, Microbiology and Molecular Biology, University of South Florida, 4202 E. Fowler Avenue, Tampa, Florida 33620-5150, U.S.A. kburtutl@uno.edu, jutley@uno.edu. Begonia campanensis Burt-Utley & Utley and B. fortunensis Burt-Utley & Utley from Panama; B. Sm. & B. G. Schub. and B. brevicyma C. DC Resumen. Begonia campanensis Burt-Utley & Utley v B. fortunensis Burt-Utley & Utley de Panama, as! como B. matudae Burt-Utley & Utley de Chiapas y B. mucronistipula C. DC. de Panama, lo que resulta en B. G. Schub. y B. brevicyma C. DC. con ella. " IUCN Red List, Mexico, Oaxaca, Panama. Neotropics by 600 or more species that are their ranges, occurring in Costa Rica and Panama forests and thorn-scrub vegetation at elevations from sea level to over 3300 m. Almost one third of the and are most abundant in western Mexico. In Flora species treated are tuberous (Burt-Utley & McVaugh, in Mexico (37 spp.) (Burt-Utley, pers. obs.). Unlike In both Mexico and Central America, many Begonia capsules and seeds that lack apparent adaptations for dispersal (Burt-Utley. 1985). Based on morpho¬ logical characters, there are two of at least 16 groups America (92 spp.). i spp.) (Burt-Utley & Utley, 1990), and section Weilbachia (Klotzsch & Oerst.) A. DC. (25 spp.) (Burt-Utley & Utley, 1999). Within the area doi: 10.3417/2011017 Novon 21: 393-401. Published on 29 December 2011. The it is i (5-11 X 6-9 vs. 9-13[- !h both species have flowers of B. to spiral when dry (Burt-Utley, 1985). i mm. (0.8-)l-2.2 cm r;1“ : 5-6.5 cm when not f t broadly elliptic with one of the pairs ! bilobed or trilohed distally. s (7.5-)9-12(— 17) mm. con- lliptic, 6-7.5 X 4-4.5 mm; known only from the type localism the Sierra Madre 401 Transfer of Four Species of Scabiosa to Lomelosia (Dipsacaceae) Universita degli Studi di Napoli Federico II, via Foria, 223, 1-80139 Napoli, Italy. a (Reck f.) P. Caputo & Del Guacchio, L. i (Rech. f.) P. Caputo & Del G :.) P. Caputo & Del Guacchio, L. . & Buhse) P. Caputo & Del i. f.) P. Caputo & are part of the epicalyx tube, not of the corona); and After the studies of Verlaque (1984, 1985, 1986a, 1986b) and Devesa (1984), which demonstrated a Adans.; Haller (1768) l. & Schult. Quer) was s h as Lopez .^Later, Sojak (1987) transferred' sevlal W Rechinger (1989) and Lack ami Rechinger (1991), expanded into a more or less membranous rim or Southwest Asia, did mrt adopt^ the nomenclatural ate epicalyx and lack of caly; und the same time, various f r (1824) and Hock (1891), p 403 and Pterocephalus (L6pez Gcmzales, 1987; Mayer & & Buhse) P. Caputo Imp. Naturalistes Moseou 12: 112-113. 1860. [1991: 48]. G-BOIS. G A New Variety of Musa itinerans (Musaceae) in Taiwan Hui-Lung Chm 2011. Volume 21, Number 2011 Chiu et al. 407 Volume 21, Number 4 2011 Chiu et al. Musa (Musaceae) in Taiwan 409 S133047-5/5 was determined by Chiu et al. (2010). the pericarps may be light initially and then darken with maturity. The principal difference between 410 Volume 21, Number 4 2011 Chiu et al. Musa (Musaceae) in Taiwan Plant height Leaf habit light-green the bract B to 11 hands, 8 to 12 fruits per hand on average 7 cm, straight and ridged lifting two bracts at a time, 9 hands, 15 fruits per hand pale green & X. J. Ge by Hakkinen et al. (2010), with pale green Pamtypes. TAIWAN. Hail [Yilan] Co.: Tatung Town¬ ship, Yingshih, #202 logging track 4.2 km, male flowers, 2 Sep. 2010, H. L. Chiu 12 (K, MO, TAI, TI, TNM), #202 Chiu 14 (KUN, TNM). Iey to the Species of Musa and Variations of M. itinesans i Taiwan 412 Novae Gesneriaceae Neotropicarum XVII: New Combinations and Typifications John L. Clark,1'2 Laurence E. Skog,2 and Fred R, Barrie? 1 Department of Biological Sciences, The University of Alabama, Box 870345, Tuscaloosa, Alabama 35487-0345, U.S.A. jlc@ua.edu 2 Department of Botany, MRC-166, National Museum of Natural History. Smithsonian Institution, P.0. Box 37012, Washington, DC 20013-7012, U.S.A. skogl@si.edu 3Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166, U.S.A. Corresponding address: Department of Botany, Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, Illinois 60605, U.S.A. flrarrie@fieldmuseum.org Abstract. Fieldwork and current research projects in Central and South America on the Gesneriaceae The application of the name C. serrata (Klotzsch ex without a Latin diagnosis, is here validly published as R. bracteata J. L. Clark & L. E. Skog. Resumen. El trabajo de campo en Gesneriaceae Ecuador, se transfiere de Dalbergaria Tussac. C. angustata (Wiehler) L. E. Skog. Se reconoce a como R. bracteata J. L. Clark & L. E. Skog. Recent fieldwork in Central and South America Wiehler, Gesneriana 1: 84. fig. 28. 1995. TYPE: Ecuador. El Oro: along rd. from Loja to Santa Rosa, 20 km past Pinas, 17 Apr. 1986, H. Wiehler & GRF Expedition 8652 (holotype. QCNE; isotypes, B, E, K, MO, NY, QCA, SEL, US). Wiehler & GRF Expedition 8652) that were "distrib- Foundation in the 1980s. This species was treated as Boggan. L. E. Skog & Roalson during a recent al., 2008). It was noted by Boggan et al. that their further studies could well indicate that the Ecuador- Clark et al. 11934 [UNA, US, USM]"and J. L. Clark et al, 11910 [BRIT, K, MO, NY, SEL, UNA, US, doi: 10.3417/2011002 Novon 21: 413-423. Published on 29 December 2011. 414 Volume 21, Number 4 2011 Clark etal. 415 Novae Gesneriaceae Neotropicarum XVII History [NMNH] Imaging). '' ? 416 Novon et al. 11910 (BRIT, K, MO, NY, SEL, UNA, US, USM). 1 Volume 21, Number 4 2011 Clark etal. 417 Novae Gesneriaceae Neotropicarum XVII QCNE, US), B from .]. L Clark, F. Nicolalde * R. Hall 7386 (MO, QCNE, SEL, UNA, US). ' P ' ' 418 Novon yellow corolla. Images A from field collection /. L Clark et al. /493 (COL, NY, QCA, QCNE, LIS), B from /. L Clark 880/ (MO, QCNE, SEL, UNA, US), C from J. L. Clark 10122 (MO). Volume 21, Number 2011 Clark et al. XVII 420 Novon Volume 21, Number 2011 Clark et al. XVII 422 Novon equal or subequal in size and the fruit, as in all (1858) indicated that he took the name, and possibly which Hanstein (1865: 390) gives as Warscewicz 19, “Costa Rica et Veragua” or “Veragua.” Thus it is not A New Name in Drymonia 2: 114. fig. 32D. 1977. non Drymonia dressleri Wiehler. Selbyana 5: 80. fig. 2C 1978. TYPE: Panama. Code: hills N of El Valle de Anton. 800 m, 17 Jan 1973. R. L. Dressier 4258 (holotype, SEL; isotype. MO). recent phylogenetic results (Clark et al.. 2006) that (Chiriquf, Code. Comarca Kuna Yala. Darien. Panama, and Veraguas) and Costa Rica (Linton). texture are features that D. ovatifolia shares with D. presented in Clark et al. (2006). Wiehler (1977) first author of the holotype (Dressier 4258) suggest that the sagittate anthers are in an early stage of described in Clark et al. (2006) as an intermediate pore stage that later develops into longitudinal Validation and New Combination of a Name in Resia F. de Jesus, BioLlania (Ed! Espec.) 6: 520. (L. E. Skog & F. de Jesus) Fern. Alonso. Revista Acad. Colomb. Ci. Exact 30: 175. 2006. nom. inval. TYPE: Colombia. Santander: Mpio. Cha- rala. 25 June 1993. X. Londono & L. P. Kvist 812 (holotype. COL; isotypes. AAU. HUA. K, MO. TULV. US. VEN). and Venezuela. Skog and de Jesus (1997) proposed a include a Latin description, therefore the name was not validly published (McNeill et al.. 2006). cies name, published by Fernandez -Alonso (2006). (Wiehler. 1983: Skog & de Jesus. 1997). All members of Napeantheae are characterized by the Clark et al. XVII Hawksworth, K. Marhold, D. H. Nicolson, J. Prado, P. C. Silva, J. E. Skog, J. H. Wiersema & N. J. Turland clature (Vienna Code). Regnum Veg. 146. Oersted, A. S. 1858. Centralamerieas Gesneraeeer, et Naturvidensk. Math. Afd. Ser. 5. Skog, L. E. & F. F. de Jesus. 1997. A review of Resia (Gesneriaeeae). BioLlania, Ed. Espec. 6: 515-525. Nogopterium, a New Name for the Genus Pterogonium (Musci, Leucodontaceae) Marshall R. Crosby Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166-0299, U.S.A. mardil230@skybest.com William R. Buck The New York Botanical Garden, 2900 Southern Boulevard, Bronx, New York 10458-5126, U.S.A. bbuck@nybg.org & W. R. Buck, are made. Hedw. (1801) and is illegitimate (Brummitt. 2002; McNeill et al.. 2006: Art. 52.1, 2). Swartz (1798) Musci after January 1. 1801 (McNeil et al.. 2006: ( Ptengynandnim . H.)” (Anonymous, 1801: 537). Pterogonium has been incorrectly used since ca. 1855 (e.g., Schimper, 1855) for a genus of Leuco- Crosby et al., 2000). No generic name is available for under Article 32.1(d) of the Vienna Code (McNeill et effectively published description.” It is an anagram of NogopLeriuin Crosby & W. R. Buck, nom. nov., pro 2, 575. 1876 TYPE: Pterigynandrum gracile Hedw. [= Nogopterium gracile (Hedw.) Crosby & W. R. Buck], Naturhist. Foren. Kjpbenhavn, ser. 4, 1: 127. 1879. TYPE: [Brazil:] “Prope Novo Friburgo ad 2. NogopLeriuin gracile (Hedw.) Crosby & W. R. gracile Hedw.. Sp. Muse. Frond. 80. 1801. Hypnum gracile (Hedw.) With., Syst. Arr. Brit. PL, ed. 4, 3: 842. 1801. Grimmia gracilis (Hedw.) F. Weber & D. Mohr. Index Mus. PI. Crypt. [2]. 1803. Pterogonium gracile (Hedw.) Sm., Engl. Bot. 16: 1085. 1803. Maschalocarpus gracilis (Hedw.) Spreng.. Syst. Veg. 4(1): 158. 1827. Leptohymenium gracile (Hedw.) Hueb- ener, Muscol. Germ. 554. 1833. Anomodon 46. 1840. Neckera gracilis (Hedw.) Mull. Hal., Syn. Muse. Frond. 2: 97. 1850. TYPE: [United Kingdom.] “Anglia et Scotia prope Edinburg: Geissler [1999: 430], “e Dillenii herbario.” G Schimp. (Jardin, 1875: 264) is noted from the E. Jardin coll., with an isotype seen at NY. The name (Anderson et al.. 1990). Novon 21: 424-425. Published on 29 December 2011. doi: 10.3417/2011089 Ht'rwSS:,”sr"F A New Species of Pachycarpus (Apocynaceae: Asclepiadoideae) from KwaZulu-Natal, South Africa Melissa Glen, Ashley Nicholas, and Jennifer Lamb School of Biological and Conservation Sciences, University of KwaZulu-Natal, Private Bag X54001, Durban 4000, South Africa. 205500248@ukzn.ac.za Adam Shuttleworth School of Biological and Conservation Sciences, University of KwaZulu-Natal, Private Bag X01, Scottsville, Pielemiaiilzhurg 3200, South Africa Abstract. Pachycarpus acidostehna M. Glen & species described from the KwaZulu-Natal Province of South Africa. The new taxon is differentiated from scaber (Harv.) N. E. Br. and P. asperifolius Meisn.) by IUCN Red List, Pachycarpus, South Africa. During recent fieldwork undertaken at Highflats in the South African province of KwaZulu-Natal, one of the (Harv.) N. E. Br. Analyses of morphological data. asperifolius (Figs. 1, 2). Although the distributions of all Based on these findings, it was decided that the Brown (1902, 1908) and currently comprises 44 taxa throughout Africa (Goyder, 1998). Thirty of these taxa to this region (Smith. 1988). Excluding the taxa within section Trichocodon D. M. N. Sm. (Smith, correlated diagnostic characters (Nicholas. 1999). nov. TYPE:' South Africa. KwaZulu-Natal Prove Ixopo Distr., 1 km SSE from Highflats, 3 Dec. 2007, A. Shuttleworth 38 (holotype, NU; isotype, MO). Figures 1A, C, 3. Haec species Pachycarpo scabro^ (Harv.) N. E. Br. et P. scabrid. Leaves oval, oblong and wide, 3. 8-7.9 X 2- to pale yellow, sweetly scented, 13-17 X 23-25 mm; Novon 21: 426-430. Published on 29 December 2011. doi: 10.3417/2010081 Volume 21, Number 2011 etal. ■s (photo by M. Glen [Af. Glen & W. Froneman 106, 428 etal. Arum cylindraceum subsp. pitsyllianum (Araceae), a New Taxon from Cyprus Georgias Hadjikyriakou Antifonitis str. 10, CY-4651 Trachoni, Cyprus, alakati@cytanet.com. cy Rolf Hand and Guilhem Mansion Botanischer Garten und Botanisches Museum Berlin-Dahlem, Freie Universitat Berlin, Konigin-Luise-Str. 6-8, D-14195 Berlin, Germany, r.hand@bgbm.org; g.mansion@bgbm.org Abstract. Arum cylindraceum Gasp, subsp. pitsyl- connectives that are orange. Key words: Araceae, Arum, Cyprus, IUCN Red List. According to Boyce (1993, 2006), the genus Arum megobrebi Lobin, M. Neumann. Bogner & P. C. infragenera (cf. Boyce. 1993. 1994. 2006) and (section Arum), A. sintenisii (Engl.) P. C. Boyce Dioscoridea subsection Tenuifila (Engl.) P. C. Boyce), subsect. Alpina P. C. Boyce. Recent molecular 2008; Espmdola et al.. 2010; Linz et al.. 2010), In 1996, Amm specimens were collected by the first author from Madari Peak in Cyprus, and the of the plant (one showing the open spathe with spadix, and one showing the leaf blade) were sent to Peter Boyce (pers. comm., 2004), who confirmed the Mediterranean) — the nearest population in the Med- populations in continental West Turkey.” Since then, accepted by Boyce (1993). were later classified Boyce (2006: 134) stated that “without a shadow of Bedalov & Ktipfer. 2005. for taxonomic concepts), but a recent molecular study by Linz et al. (2010: The study by Espmdola et al. (2010: 27) indicated Material and Methods of the new taxon. Plants collected in the field were doi: 10.3417/2011007 Novon 21: 431-436. Published on 29 December 2011. et al. et al. Lectotypification and Reinstatement of Stachytarpheta friedrichsthalii (Verbenaceae), with Notes on the Lectotypification of S. indica Barry E. Hammel and Michael H. Grayum Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166-0299, U.S.A. barry.hammel@mobot.org; michael.grayum@mobot.org Abstract. Certain material of Stachytarpheta Vahl of the latter name ( Fendler 219. MO; Friedrichsthal in synonymy. One of the two sheets of Friedrichsthal negative 34320 — is chosen as the lectotype. Contrary consequent synonymy of S. angustifolia (Mill.) Vahl correct name for some African material that has been friedrichsthalii has been found among Central treatment of Verbenaceae (in the traditional sense) for margined leaves and 4-toothed calyces). It should be calyx follows Pool (2001) and varies slightly front that used by Atkins (2005), who described the calyx of S. indica (as S. angustifolia (Mill.) Vahl) as abaxially 2- proposed by Moldenke and Moldenke (1983: 266) description” (Atkins, 2005: 193). However, the late marginatis”; Linnaeus, 1762: 27), albeit discor- inconsequential with regard to lectotypification. Conversely, the lectotype of V. indica (the larger spicis longissintus camosis nudis. foliis lanceolato- rnust be accepted. Moldenke and Moldenke (1983)! Fernandes (1984), and Atkins (2005) all acknowl¬ edged that this lectotypification relegates S. angus- consequence. Atkins (2005: 193), nevertheless, angustifolia and the other, with crenate and non- as V. indica by Linnaeus in 1762), which she says second taxon may not be restricted to Africa at all, I buck. sensuVool, 2001; Atkins, 2005) has corn/to fight, numerous Costa Rican collections of S. calderonii moist lowlands of the north to central Pacific slope. (see key below) follows that in current use in Central doi: 10.3417/2011019 Novon 21: 437-439. Published on 29 December 2011. Changes to Publication Requirements Made at the XVIII International Botanical Congress in Melbourne — What Does e-Publication Mean for You? doi: 10.3417/2011072 441 [) or ers final, in « (ISBN). Publication is not ef cation of new names at a pul placing of names in collection the public, by the issue of n produced at a later date is not eff, 30.4. For the purpose of this Article, it The content of a e purpose of obtaining a degree is published unless it includes an a as far as is praetieal (see a 11 d printed matter of any kind, it in terms of number of copies is not ol single hard copy papers as should the volume be great. scientific names, which continue to be Latin or :al.. 2011). is that all new names Thirteen New Species of Neotropical Yiscaceae (. Dendrophthora and Phoradendron ) Victoria, BC V8W 3 Volume 21, Number 4 2011 Kuijt New Species of Neotropical Viscaceae 445 nearly acicular basal cataphylls to 4 mm, in median 446 Novon position, 1-1.5 cm above the base, basal cataphylls acute, base tapering into indistinct petiole ca. 1 mm; 447 448 Novon and the small, few-flowered inflorescences. The new from Venezuela (Kuijt. 2003a). but. aside from the fertile bract (vs. one and distally located) in a staminate inflorescences may well be more florifer- Volume 21, Number 4 2011 Kuijt New Species of Neotropical Viscaceae 449 San Martin, Nuevo Peru", 5°22'S, 78°30'W, 900 m, 14 May 1996 (c? fls.), R. & A. Vdsquez 20837 (holotype, UC; isotype, MO-4782656). Figure 5. 450 Novon nearly halfway to the nearest foliar node. Leaf blades obscure veins, the middle one not reaching the apex. Volume 21, Number 4 2011 Kuijt New Species of Neotropical Viscaceae 451 ly short, to 1.5 mm. followed by 5 to 7 fertile in biseriate rows. Pistillate plants not known. Deficient, according to IUCN criteria (2001). The related to the more northern D. costaricensis Urb. The 6. Dendrophthora verrucosa Kuijt. sp. nov. TYPE: Mar. 1998 ( c? fls.)". //. van der Werff, B. Gray, R. Vasquez & R. Rojas 14937 (holotype. UC; isotype. MO not seen). Figure 6. basal cataphylls as 1 pair. 5-10 mm above the base. lateral veins: leaf margin smooth, translucent when drophthora verrucosa is assessed as DD or Data Deficient, according to IUCN criteria (2001). and is Kuijt (Kuijt. 1990). but D. verrucosa differs in being distinctly verrucose rather than glabrous, in having terete rather than flattened and grooved intemodes. localities of D. dimorpha and D. verrucosa are slightly vie. of Poblado Los Llanos. 5°6'16"S. 78°51'11"W. 1875 m, 12 Oct. 2006 (? fl.), J. Perea & V. Flores 2831 (holotype. UC; isotypes. AMAZ not seen. HUT not seen. MO not seen. MOL not seen. USM not seen). Figure 7. to 5 X 1.2 cm. strongly quadrangular to 4-winged. followed by ca. 6 fertile intemodes each 8-10 mm Deficient, according to IUCN criteria (2001). somewhat winged intemodes. Unfortunately, the type 452 Novon variable (Kuijt. 2003a). and dioecv in the present Jose del Rio Chirinos (Rio rnira flores). 5°12'S. 78°46'W. 600-700 m, 16 Apr. 1996. J. Campos 7. Phoradendron datum Kuijt. —A. Fertile habit. — B. Inflorescence. A, B drawn from the holotype Perea & Flores 454 Novon 456 Novon Para-types. VENEZUELA. Miranda: Arboretum Es- 291 (UC), 292 ( U C ) ; on Zanthoxyhim dliatum Engl., Lopez 293 (UC). TYPE: Pent. Amazonas: Rodriguez de Mendoza, Mariscal Benavides. Izcuchaca. 6°19'40"S, 77°31'5"W, 1880 m, 29 Aug. 1998 (c? fls.). Volume 21, Number 4 2011 Kuijt New Species of Neotropical Viscaceae 457 R. Vasquez & J. Campos 25306 (holotype, UC; isotype, MO-1510519). Figure 11. nearly 1 cm below foliar nodes; basal cataphylls 1 : 1 1 . Phoradendron nickrentianum Kuijt. —A. Fertile habit. — B. Inflorescence. A, B drawn from the holotype Vdsque: & os 25306 (UC). Ilf r 460 Novon holotyp l~QuLpeYwZm 2496(1 113, fig. Volume 21, Number 2011 Two New Species of Oryctanthus (Loranthaceae) from Colombia and French Guiana 2011. 464 Novon Notes on Tetrorchidium (Euphorbiaceae) in Panama 470 Literature Cited Correa, M. D„ C. Galdames & M. S. de Stapf. 2004. Strychnos puberula (Loganiaceae), a New Species from Panama Gordon McPherson Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166-0299, U.S.A. gordon.mcpherson@mobot.org Abstract. Strychnos puberula McPherson (Logania- (Aubl.) Mart, in its indument of minute hairs, lack of short-stipitate, relatively thick-walled fruit. Key words: IUCN Red List, Loganiaceae, Pana- not be matched to any of the 11 species listed in the (Correa et al., 2004) and represent an undescribed 2-4.2 cm vs. ca. 15 X 8 cm) and 1 -seeded fruits "(vs. Strychnos puberula McPherson, sp. nov. TYPE: 080°41'05"W, lowland forest, 150 m, 16 Apr. 2009, collected with J.-Y. Serein, G. McPherson 21021 (holotype. PMA; isotype, MO). Figure 1. Woody vine in canopy trees; branchlets slightly cellate: tendrils (uncoiled) ca. 7-8 cm. appressed- 10.2 X 2-4.2 cm, base obtuse, apex acuminate. 5- Novon 21: 472-474. Published on 29 December 2011. plinerved, the basal pair of secondary veins arising at arising (5— )8— 12 mm distally; midrib slightly im- adaxially, more evidently raised abaxially; the adaxial surface; raised portion of stem at base of petiole 1.5- 2 mm, appressed-puberulent; petiole 4-6 mm, channeled, appressed-puberulent. Inflorescences ax¬ illary, raceme-like, of 5 to 9 flowers, the axis (2— )5— ca. 2 mm; pedicels 1-3 mm in flower. 2-4 mm in fruit. Flowers 4-merous; calyx 1-1. 5(— 2) mm, the abaxially; corolla white, ca. 6.5 mm, tube 3-3.5 submedially affixed, ca. 0.8 mm; ovary 1 mm, glabrous; style 6 mm. glabrous. Fruit pale yellow (not fully mature), subspherical. 14-20 mm diarn. status of DD or Data Deficient (IUCN, 2001)' size (4—7 X 1. 5-3.1 cm in the latter species) and doi: 10.3417/2010095 Volume 21, Number 2011 473 474 in G. Davidse, M. Sousa S„ Knapp & F. Chiaiig (editors). A New Genus of Podostemaceae from Venezuela C. Thomas Philbrick and Jacqueline Malecki Department of Biological and Environmental Sciences, Western Connecticut State University, Danbury, Connecticut 06810, L.S.A. philbricktObwcsu.edu Nicholas P. Tippery Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269-3043, U.S.A. Current address: Department of Biological Sciences, University of Wisconsin- Whitewater, 800 West Main Street, Whitewater, Wisconsin 53190, U.S.A. Hannah I. Stevens GIS Program Manager, New York Botanical Garden, 200th Street and Kazimiroff Boulevard, Bronx, New York 10458, U.S.A. illustrated and described. The new species is Castelnavia Tul. & Wedd.. Oserva Tul. & Wedd.. Castelnavia Tul. & Wedd.. Oserva Tul. & Wedd.. Noveloa C. T. Philbrick y Rhyncholads Tul., as! corno Resumen. Se ilustran y describen un genero Autana surge de una continuidad de los margenes de la hoja. donde los estambres caedizos estaban unidos. se doi: 10.3417/2010051 22 species in the country. Philbrick et al. (2010) listed the same genera, but somewhat more (28) Venezuelan Guayana alone. Six of the genera occur in the Weddellenoideae ( Weddellina Tul.) or Tristichoideae (Tristicha Thouars) (Royen, 1951, 1953, 1954). The current contribution adds an were collected that did not correspond to any of the studies (ITS, rbcL, trnL) of Neotropical Podostema- al., 2011, listed as “Autana,” see below). Conse- Novon 21: 475-480. Published on 29 December 2011. 476 2011 478 Novon Volume 21, Number 4 Philbrick et al. 479 201 1 Autana (Podostemaceae), a New Genus from Venezuela Jan. 1949, Maguire & Politi 28400 (NY); Puerto Ayacucho, Rfo Cataniapo, ca. 15 km E of main rd., 5.5548°N, 67.4856°W, 9 Jan. 2006, Philbrick, Novelo & Lasso 5878 (CAR, MO, VEN, WCSII); Puerto Ayacucho, Rio Cata- 67.5937°W, 9 Jan. 2006, Philbrick, Novell & Lasso 5875 (CAR, MO, VEN, WCSU); Canyo Marieta, ca. 1 hr. by boat upstream oi coniluence with Rfo Ventuari, 5.1687°N, 66.5334°W, 8 Mar. 2009, Philbrick, Fort & Perre: 6318 (VEN, WCSU). Acknowledgments. We thank Albino Luna for National Science Foundation Grant DEB -0444589 Philbrick et al 5862 (VEN); B froni Philbrick et al. 5867 (WCSU); D-G from Philbrick et al. 5875 (WCSU). ” Gouania tiliifolia: The Correct Name for Gouania scandens Peter B. Phillipson Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166-0299, U.S.A.; and Museum national d'Histoire naturelle, Departement Systematique et Evolution, UMR 7205, Case Postale 39, 57 rue Cuvier, F-75231 Paris CEDEX 05, France, peter.phillipson@mobot.org Martin W. Callmander Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166-0299, U.S.A.; and Conservatoire et Jardin botaniques de la ville de Geneve, ch. de Llmperatrice 1, 1292 Chambesy, Switzerland Sven Buerki Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3DS, United Kingdom Abstract. In a recent revision of Gouania Jacq. ex l iil.i Buerki, Phillipson & Callm. is provided for incluant les autres ties de l’Ocean Indien occidental, Cette erreur est corrigee et la nouvelle combinaison G. tiliifolia subsp. glandulosa (Boivin ex Tul.) Jacq. (Rhamnaceae) for Madagascar and the other western Indian Ocean islands (Buerki et al., 2011), a 1783 and 1808. contained descriptions of five tiliifolia Lam, occur in the region concerned. After (Britten, 1906; Stafleu & Cowan, 1979) we estab¬ lished that the correct date of publication was 1789 manuscript. Unfortunately, we failed to notice that name G. tiliifolia to have priority over the name G. published in 1791 (based on Retinarui scandons Phillipson & Callm. (based on G. glandulosa Boivin 1789, as “ ' tiliaefoliaf non Gouania tiliifolia Rottb. ex DC., 1825. nee Gouania tiliifolia TYPE: [Reunion Island], lie Bourbom s.d. (fl. doi: 10.3417/2011078 Novon 21: 481-482. Published on 29 December 2011. Una Especie Nueva de Combretum (Combretaceae, seccion Combretum) de la Costa del Occidente de Mexico 484 Novon A New Species of Solanum (Solanaceae) from the Highlands of Central Brazil u CEP 70.670-350 E Caroly n E. B. Proenga A New Species of Iochroma (Solanaceae) from Ecuador Stacey D. Smith Department of Biology, Duke University, Box 90338, Durham, North Carolina 27708, U.S.A. Current address: School of Biological Sciences, Manter Hall, University of Nebraska, Lincoln, Nebraska 68588-0118, U.S.A. sdsmith@unl.edu Segundo Leiva Gonzalez Museo de Historia Natural, Universidad Privada Antenor Orrego, Av. America Sue 3 1 15-1 d>. Monserrate Casilla Postal 1075, Trujillo, Peru. Segundo_Leiva@hotmail.com Abstract. Iochroma baumii S. D. Sm. & S. Leiva distribution and many floral features with its closest S. Leiva (Solanaceae) de los bosques nublados del y caracterfsticas florales con las especies hermanas: en ramas maduras. se distingue facilmente de las otras especies de Iochroma. Key w from Colombia to Pent (Shaw, 1998; Hunziker, 2001; Smith & Baum, 2006, Leiva, 2009). A member of Physaleae Miers (Olmstead et al., 2008), the genus is (Shaw, 1998). Iochroma species are most common in 2008). stead et al. (1999. 2008). Most of the species of designated with the letters A, C, L, and F (Smith & Baum, 2006; Table 1). There is an additional small clade of five Iochroma species (the U clade) that trees and more distant in others (Smith & Baum, Datura L. (Smith & Baum, 2006; Olmstead et al., 2008; Table 1). 12 new species have been described since 1995 (Leiva, 1995, 2005, 2006, 2007, 2009; Leiva et al., 1998; Leiva & Quipuscoa, 1998; Lezama et al.. Smith et al.. 2008: Table 1) as interspecific & Baum. 2007). Still, the extent of contact and largely unknown. In addition to the taxa listed in doi: 10.3417/2010061 Novon 21: 491-495. Published on 29 December 2011. 492 Novon (Shaw, 1998), we list the 12 spe Smith and Baum (2 Country abbreviations are as follows: Argentina (ARC), Bolivia (BOL), Colombia (COL), Ecuador (ECU), Peru (PER); SDS and SLG ellipticum (Hook, f.) Hunz. (Lindt.) M. I mifoliu, until) Mier exD’Arcy Hybrid (L. ) Schltdl. or other member of A clade1 more closely related to Dunalia Kunth. species than to lochroma Benth1 to Eiiolarynx (Hunz.) the cloud forests near Papallacta in northern Ecuador. Phylogenetic analysis places this taxon in fachsioides (Bonpl.) Miers) and another blue-flowered 2811 , QCNE, WIS). Fig- 4.8 X 2-2.8 r al.. IUCN Red List c of Peru), and thus the size of the populations is not 494 Novon :ktE£ UBIA. Caldas: Hoya del rio Otun, •cams 23329 (F). ECUADOR. Napo: 00°13'S, 78°02'WN/. L. Clark 3587 ■rberi MO, QCNE); carr. Rubiacearum Americanarum Magna Hama Pars XXIX: Overview of the Neotropical Genus Schizocalyx (Condamineeae) and Description of Two New Species Charlotte M. Taylor Missouri Botanical Garden, P.0. Box 299, St. Louis, Missouri 63166, U.S.A. charlotte.taylor@mobot.org David A. Neill Fundacion Jatun Sacha, Casilla Postal 17-12-867, Quito, Ecuador, davidneill53@gmail.com Roy E. Gereau Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166, U.S.A. roy.gereau@mobot.org in bud, capsules that are loculicidal across the apical part, and numerous flattened to angled seeds, as eastern Andean foothills of central Peru and for S. cuspidatus (A. St.-Hil.) Kainul. & B Bremer and S. multiflorus (Hook, f.) Kainul. & B. Bremer. Resumen. El genero Schhocalvx Wedd. original- lo cual estaban clasificadas erroneamente. y dos especies transferidas de Phi-topis Hook, f., lo cual es cidales en la parte apical y semillas aplanadas o anguladas. corno se detalle en una Central a traves de Sudamerica occidental hasta Brasil central y suroriental. Las nueve especies and S. multiflorus (Hook, f.) Kainul. & B. Bremer. Condamineeae, Costa Rica, Ecuador, IUCN Red List, The genus Schhocalyx Wedd. ( Weddell, 1854) was Novon 21: 496-507. Published on 29 December 2011. doi: 10.3417/2008095 Volume 21, Number 4 2011 Taylor et al. New Species of Schizocalyx 497 neae also by Robbrecht (1988), but shortly afterward (1991) transferred Schizocalyx to their newly de- the Calycophylleae and considered it to be related in the Rondeletieae (Robbrecht, 1988). Delprete tieae (Delprete, 1999). Later Rova et al. (2002) studied the various genera that have been included in tieae, and their results showed that all of these genera (2010), using both molecular and morphological particularly closely related. Kainulainen et al. al.’s classification (2010). These two genera were Bathysa (Standley. 1931a: 259) versus convolute in Schizocalyx (Standley. 1930: 171); however, this Krause (1908), who described B. peruviana K. Standley (1931a, 1936), who described and keyed B. Burger and Taylor (1993), who incorrectly described septicidally for more than half their length in Bathysa capsule in Schizocalyx (Kainulainen et al., 2010). sometimes overlooked (e.g., Delprete, 1997). years later, P. sterculioides Standi, of Peru (Standley, Phitopis was provided by Standley (1936). The name irregularly lobed calyx limbs. However, these genera (1995) excluded Phitopis from the tribe Hippotideae. Rondeletieae together with Bathysa, as did Standley (Robbrecht, 1988; Rova et al., 2002; Kainulainen et workers (e.g., Robbrecht, 1988), but Delprete (1999) Taylor et al. ,■ (Dwyer) C. M. Taylor (Taylor et al., : name B. multiflora L. 0. Williams was (Taylor & Pool, 1993): this name is a australis (A. St.-Hil.) Hook. f. ex K. S Petitions Working Group, 2008). s are not being submitted to IUCN 73. 1854, nom. cons. bracteosus Wedd. SRt&riiEr Taylor et al. d Peru (Taylor* Pool 1993), w Discussion. The collection Spruce 4319 was t B, versus one of tl be seen by visiting K. Of the two specimens 4319 at K. where Hooker worked, the r. J. Bot. 97(12): 1976. 504 Novon playa, 1798, J. J. Tafalla s.n. (holotype, Bf, F photo neg. #33 at MO; isotypes, F not seen, MA not seen, F photo neg. #29653 at MO). Romero, 1993; Taylor & Pool, 1993), in wet forests at 290-1700 m. This species has also been reported from Ecuador (Neill & Ulloa, 2011) based on an tioned by Krause (1908), who contrasted his new “B. obovata (Ruiz & Pavon) K. Schumann,” which is not lectotypified here, because further documen- Some inaccurate morphological observations made ment and somewhat variable flower and fruit size. Bremer, ^ Amer. J. Bot. 97(12): 1976. 2010. Bot. Vereins Prov. Brandenburg 50: 96. 1909. 1903, E. Ule 6768 (holotype, Bf, F photo neg. #32 at MO). (Taylor & Pool, 1993; Taylor, 1999), in wet forests at 200-1600 m. dispersal of the seeds (Figure 1A-C). There also Peila, L Tsamajain & M. Roca 8034 (MO). Sail Martin: SdiLheJv. & M. Dillon 8953 (F, MO). ^ Bretiie/ Amer. J. Bot. 97(12): 1976. 2010. Field Mus. Nat. Hist.. Bot. Ser. 8: 341. 1931. TYPE: Peru. Junm: dense forest, Pichis Trail, Enenas, 1600-1900 m, 30 June-2 July 1929, E. P. Killip & A. C. Smith 25747 (holotype, F- 607646 F photo neg. #44915 at MO). & Pool, 1993; Neill & Ulloa, 2011), where it is found in wet premontane forests at 1100-1800 m. It has talline rocks (e.g.. D. Neill & Shuar Conservation Young & G. Sullivan 747, MO). El Paujil, lowland rainforest, 10°10'24"S, 75°15'49"W. 500 m. H. van der Werff, R. Vdsquez & R. Francis 19999 (holohpe. USM; isotype, MO-04824519). Figure 2. Taylor et al. Hoy a ignorata (Apocynaceae, Asclepiadoideae): An Overlooked Species Widely Distributed across Southeast Asia Tran The Bach Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam Joo-Hwan Kim and Dong-Kap Kim Department of Life Science, Kyungwon University, 65 Bokjeong-dong, Seongnam, Gyeonggi-do 461-701, Korea Joongku Lee Korea Research Institute of Bioscience and Biotechnology, Eoeun-Dong 52, Yuseong-Gu, Daejeon 305-333, Korea Bui Thu Ha Hanoi National University of Education, 136 Xuan Thuy Street, Cau Giay District, Hanoi, Vietnam Nadhanielle Simonsson Juhonewe P.0. Box 1-524, Ukarumpa, Eastern Highlands Province 444, Papua New Guinea Michele Rodda Singapore Botanic Gardens Herbarium, 1 Cluny Road, 259569 Singapore. Author for correspondence: rodda.michele@gmail.com ceae. Asclepiadoideae) is described and illustrated Southeast Asia, does not seem to be related to any the genus Hoya, are discussed in light of the recent following the results of molecular phylogenies and morph ological studies. Key words: Apocynaceae, Hoya , IUCN Red List, Malaysia. Sabah. Thailand. Vietnam. Pacific Islands (Li et al., 1995). The plants are often 1912; Ho, 1993; Li et al., 1995; Tran, 2005, Rodda Peninsular Malaysia contains 26 Hoya species (Rintz, 1978, Kiew, 1989). Thailand is rich in Hoya species, with ca. 40 taxa (Thaithong, 2001); Borneo is still little explored (Forster et al., 1998), but recent description (Rodda & Nyhuus. 2009: Rodda et al.. 2011; Rodda & Simonsson, 2011a, 2011b). Hoya has have been greatly expanded (Wanntorp et al., 2006a, 2006b, 2011; Wanntorp & Forster, 2007, 2009; Wanntorp & Meve, 2011). Rare morphologies such as Rodda & Simonsson. 2011b) and extremely minute describe the new species as //. ignorata T. B. Tran, Novon 21: 508-514. Published on 29 December 2011. doi: 10.3417/2010068 Tran et al. Hoya (Apoi 510 Novon paratypes Simonsson & Somadee NS10-004, NSlO-OOt (SING). Photo by N. Simonsson. — B. Mature specimen, drawn from the Sabah, Malaysia (photos'" by M. Rodda) Volume 21, Number 2011 Tran et al. Hoya (Apoi 511 512 Novon terminal appendages (Omlor, 1996). Within Marsde- seeds are comose and spindle-shaped and lack outer margin (Schill & Dannenbaum. 1984). This broadly margined, smooth seeds (Omlor. 1996). 2006a. 2006b. 2011) it became apparent that Hoya Forst., Liddle & I. M. Liddle, and Micholitda N. E. Br., and the two species of Clemensiella (Schltr.) Schltr. (Wanntorp & Forster. 2007; Wanntorp & Kunze, 2009; Meve et al.. 2010; Wanntorp & Meve. urceolate corolla, and in Micholitda . the small. different genera. Similarly. C. mariae (Schltr.) Schltr. despite bearing fleshy coronas with valvate lobes and narrow retinacula, and long, ribbon-shaped cau- this habit have recently been described (Rodda et al.. in prep.; Rodda & Simonsson, 2011b). The new taxon comose. spindle-shaped and lack differentiated H. m ari ae (Schltr.) L. Wanntorp & Meve and //. (Wanntorp & Meve. 2011). deae from Marsdenieae (Bruyns & Forster. 1991) and Only H. ininutiflora Rodda & Simonsson (2010) Paratypes. MALAYSIA. Sabah: Nabawan, 300 m.s.m., 13 June 2011, M. Rodda & L. Gokusing MR11-056 (SING). Terengganu: kennman, Bukit Kajang, 150 m.s.m., 6 Nov. 1935, E. H. J. Corner 30279 (SING). THAILAND. Nakhon Si Thamniarat: Khao Luang, 800 m.s.m., 5 May 2010, N. Simonsson & S. Somadee NS10-004 (SING), 810 m.s.m., 6 May 2010, N. Simonsson & S. Somadee NS10-007 (SING). Tran et al. Hoya (Apoi 513 514 Lectotypification of Three Species in the Fern Genus Pteris (Pteridaceae) from China Yang Dong-Mei Xing Fu-Wu and Wang Fa-Guo* P. esquirolii H. Christ, and P. henryi H. Christ. Distribution for P. dalhousieae, which is not found in doi: 10.3417/2010094 21: 515-516. 2011. 516 Author Index: Novon Yol. 21] An, M.-T., H.-H. Zhang, Q. Lin & T.-L. Wei. Arisaema Attanayake. A. M. A. S„ I. M. Turner & R. M. K. Saunders. Tozzi. 331-337 Bacchetta, G., S. Brullo, T. Cusma Velari, L. Feoli Chiapella Bai, D.-Q. see Shao et al. 256-261 Barrie, F. R. see Clark et al. 413-423 chilus tetraptenis to Odontochilus (Orehidaeeae), 20-23 Bolin, J. F., R. D. Bray & L. J. Musselman. A New Species of Lebanon, 295-298 Bray, R. D. see Bolin et al. 295-298 Brullo, S. see Bacchetta et al. 4—19 Buck, W. R. see Crosby & Buck. 424—425 Buerki, S. see Phillipson et al. 481-482 Burt-Utley, K. & J. F. Utley. New Species and Notes on 401 Bush, C. M. see Fritsch & Bush. 338-342 Callmander, M. W. see Phillipson et al. 481-482 Caputo, P. & E. Del Guacchio. Transfer of Four Species of Castro, A. F. see Rendon Sandoval et al. 483-486 Chang, Z.-Y. s, Chen, H.-F. se, Chen, H.-Q. se ■e Wang et al. 278-280 e Liu et al. 216-218 Wang et al. 380-384 > Xu et al. 154-157 > Xu et al. 285-289 Chen, L„ Z.-G. Zhang, Y. Hu, X.-W. Li & J.-Q. Li. A New from Hainan, China, 317-321 Chen, Y. T. see Wang et al. 278-280 Musa itinerans (Musaceae) in Taiwan, 405-412 tions, 413-423 " ceae) from Ecuador, 36-39 Del Guacchio, E. see Caputo & Del Guacchio. 402-404 Delgadillo, R. R. see Rendon Sandoval et al. 483—486 Delprete, P. G. see Taylor et al. 133-148 New Species from Madre de Dios, Pent, 322-325 Dogan, M. see Celep et al. 34-35 Duan, L.-D. see Lin et al. 212-215 If—' i ,1 ... 1 1. M. Longhi Wagner & T. T de Souza dries. New Pooideae, Poeae), 326-330 Fang, Z. see Yan et al. 388-391 Fritsch, P. W. & C M. Bush. A New Species of' Gtmltheria 338-342 Gereau, R. E. see Taylor et al. 118-132 Gereau, R. E. see Taylor et al. 133-148 Gereau, R. E. see Taylor & Gereau. 154-157 21(1) pp. 1-160, 21(2) pp. 161-294, 21(3) pp. 295-392, 21(4) pp. 393-528. Novon 21: 517-520. Published on 29 December 2011. 518 Novon Gereau, R. E. see Taylor et al. 496-507 Glen, M., A. Nicholas, J. Lamb & A. Shuttleworth. A New from KwaZulu-Natal, South Africa, 426-430 (Papaveraceae), 182 from the Venezuelan Guayana, 183-186 H Cyprus, 431-436 Hammel, B. see Taylor et al. 118-132 Hand, R. see Hadjikyriakou et al. 431-436 He, S.-Y., P.-T. Li, J.-Y. Lin, G. Lin & H.-L. Zeng. Hoya Hainan. China. 343-346 He, S.-Z., W.-L. Xu, Y.-Y. Wang & Q.-W. Sun. A New 187-189 Calathea (Marantaceae) from Panama, 201-211 Kim, D.-K. see Tran et al. 508-514 Kim, J.-H. see Tran et al. 508-514 Kim, Y.-D. see Shin et al. 373-374 Knapp, S„ J. McNeill & N. J. Turland. Changes to Publication Mean for You?, 440-443 (Dendrophthora and Pharadendron), 444—462 Lamb, J. see Glen et al. 426-430 Li, J.-Q. see Chen et al. 317-321 Li, P.-T. see He et al. 343-346 Li, Y.-D. see Xu et al. 285-289 Li, Z. see Ma et al. 349-353 Lima, H. C. see Mendonga Lilho et al. 73-77 Lin, G. see He et al. 343-346 Lin, J.-Y. see He et al. 343-346 Lin, Q. see An et al. 1-3 Lin, Q. & Z.-R. Yang. Validation of Two Names, Berchemia J Jia, Y. see Yu et al. 290-293 Jiang, N„ X.-M. Peng & W.-B. Yu. Valid Publication of Mexico, 192-195 Liu, X. see Ma et al. 349-353 Liu, X.-L., X.-N. Yue, Z.-Y. Chang & L.-R. Xu. A New Xinjiang, China, 216-218 Liu, Y. see Xu et al. 385-387 Longhi Wagner, H. M. see Essi et al. 326-330 Lopez. M. G.. A. L. Wulff & C. C. Xifreda. Senecio ser. ica, 347-348^ Lorence.^D. H. see Taylor et al. 133-148 Lu, S.-G. see Shao & Lu. 90-93 Volume 21, Number 4 2011 Author Index 519 M Ma, H., Q. Pan, L. Wan, Z. Li, Y. Wan & X. Liu. Musella from Sichuan, China, 349-353 Malecki, J. see Philbrick et al. 475-480 McNeill, J. see Knapp et al. 440-443 Mendonga Filho, C. V., H. C. Lima, E. R. Forni-Martins & M. Musselman, L. J. see Bolin et al. 295-298 Neill, D. A. see Taylor et al. 496-507 Nicholas, A. see Glen et al. 426—430 Oliveira, R. C. de. see Ramos et al. 368-372 Pan, B. see Xu et al. 385-387 Pan, Q. see Ma et al. 349-353 longistylus (Violaceae), 249-250 Peng, X.-M. see Jiang et al. 190-191 Pennington, T. D. see Dexter & Pennington. 322-325 Persson, C. see Taylor et al. 133-148 Philbrick, C. T., J. Malecki, N. P. Tippeiy & H. I. Stevens. A Ramos, D. M., J. F. M. Vails, R. C. de Oliveira & D. Rendon Sandoval, F. J^, R. R. Delgadillo & A. F. Castro. Una ee Taylor et al. 133-148 Secco, R. de S. see Bigio & Secco. 169-173 Semple, J. C. see Lopez Laphitz et al. 219-225 Seo, M. N. see Paula-Souza & Seo. 249-250 Shao, J.-R., M.-L. Zhou, X.-M. Zhu, D.-Z. Wang & D.-Q. Bai. 256-261 Shii, C.-T. see Chiu et al. 405-412 Shin, H., Y.-D. Kim & S.-H. Oh. A New Combination in Shui, Y.-M. see Lin et al. 212-215 Shuttleworth, A. see Glen et al. 426—430 Skog, L. E. see Clark et al. 413-423 Souza Chies, T. T. de. see Essi et al. 326-330 Stevens, H. I. see Philbrick et al. 475-480 Sun, Q.-W. see He et al. 187-189 520 Novon 118- C. Persson, "r G. Delprete & R. E. Gereau. Rubiacea- ram Americanarum Magna Hama Pars XXVIR: New Taylor, C. M., D. A. Neill & R. E. Gereau. Rubiacearum Americanarum Magna Hama Pars XXIX: Overview of Description of Two New Species, 496-507 Species from Thailand (Poaceae, Panicoideae), 149-153 Tian, X. J. see Sheng & Tian. 262-265 Tippeiy, N. P. see Philbrick et al. 475-480 Tozzi, M. G. A. see Mendonga Filho et al. 73-77 Wang, W.-T. see Wei & Wang. 281-284 Wang, Y.-Y. see He et al. 187-189 Wei, T.-L. see An et al. 1-3 Wei, Y.-G. & W.-T. Wang. Elatostema recurmramum xi, China, 281-284 Wulff, A. F. see Lopez et al. 347-348 cies Widely Distributed across Southeast Asia, 508-514 Tseng, Y.-H. see Wang et al. 278-280 Turland, N. J. see Knapp et al. 440—443 •e Burt-Utley & Utley. 393-401 Madre Oriental, Mexico, 274-277 ia, N.-H. see Vu et al. 375-379 ifreda, C. C. see Lopez et al. 347-348 W see Wang et al. 380-384 W see Yang et al. 515-516 Xu, H„ Y.-D. Li & H.-Q. Chen. A New Species of Sciaphila Xu, H., Y.-D. Li, H.-J. Yang & H.-Q. Chen. Two New Species China, 285-289 Xu, L.-R. see Liu et al. 216-218 Yan, H., Z. Fang, H. Zhang & S. Yu. Fallopio, multiflora var. China, 388-391 from China. 515-516 Yang, H.-J. see Xu et al. 285-289 Yang, T. Y. 7 Yang, Z.-R. see Lin & Yang. 71-72 Yu, N.-N, Y. Jia & J.-C. Zhao. Synonymy and Typifications in Wan, Y. see Ma et al. 349-353 Wang, C.-C. see Tseng & Wang. 270-273 Wang, C.-C., Y.-H. Tseng, Y.-T. Chen & K.-C. Chang. A New Wang, D.-Z. see Shao et al. 256-261 Wang, F.-G. see Yang et al. 515-516 •e Liu et al. 216-218 Zeng, H.-L. see He et al. 343-346 Zhang, H. see Yan et al. 388-391 Zhang, H.-H. see An et al. 1-3 Zhang, Z.-G. see Chen et al. 3 Zhao, J.-C. see Yu et al. 290-293 Zhou, M.-L. see Shao et al. 256-261 Zhu, X.-M. see Shao et al. 256-261 17-321 Subject Index: Novon Yol. 21 12 Aegiphila, 44-46 Alibertia, 133—148 ' Alibertia atlantica, 133-148 Alibertia duyeri, 133-148 Alibertia edulis, 133-148 Alibertia patinoi, 133-148 Alloplectus. 413—423 Alpuua, 270-273 sect. Alpinia subsect. Catimbium, 270-273 Alpinia zerumbet, 270-273 Amalophrllon, 413-423 Anialophyllon divaricatum, 413—423 Amalophrllon ecuadoranum, 413-423 Vmazoma. 160-173 Anoectochilus, 20-22 Anthericaceae, 36-39 Apinagia, 475-480 Apocynaceae. 343-346, 426-430, 508-514 Araceae, 1-3, 431-436 Argentin Aristolochia, 285-289 Aristolochia bambusifoha, 285-289 Aristolochia championii, 285-289 Aristolochia jianfenglingensis, 285-289 Aristolochia ledongensis, 285-289 85-289 tia, 182 Arum cylindraceum, 431-436 subsp. pitsyllianum, 431-436 \nmdmclla. 149-153 A santm, 190-191 Asclepiadoideae. 426-430. 508-514 e, 69-70 Aspidistra , 187-189 sect. Aspidistra ser. Fimbriatae, 187-189 Aspidistra fenghuangensis. 187-189 Astragalus, 216-218 Astragalus bacrlukensis , 216-218 Balsas, 196-200 Balsas guerrerensis, 196-200 Bambusoideae, 40-43 Bathysa, 133-148, 496-507 Bathysa paiiamensis, 133-148 Begonia, 393-401 Begonia hrevicynia, 393-401 Begonia campanensis, 393-401 Begonia davidsoniae, 393-401 Begonia fortunensis, 393-401 Begonia makrinii, 393-401 Begonia niatudae, 393—401 Begonia niucronistipula, 393-401 Begoniaceae. 39.3-401 Bolivia, 78-89, 94-117, 219-225, 444-462, 496-507 Borojoa panamensis, 133-148 Borojoa patinoi, 133-148 Brazil, 28-33, 73-77, 169-173, 331-337, 368-372, 487- 490, 496-507 Bnza, 326-330 Bromeliaceae. 362-367 Bryopsida, 290-293 C Calathea, 49-57, 58-65, 66-68, 201-211 1 21(1) pp. 1-160, 21(2) pp. 161-294, 21(3) pp. 295-392, 21(4) pp. 393-528. Novon 21: 521-527. Published on 29 December 2011. 522 Novon sect. Calathea, 49-57, 201-211 Calathea carlae , 201-211 Calathea chiriquensis, 201-211 Calathea fredgandersii, 201-211 Calathea recurvata , 49-57 Calathea tarrazuensis , 49-57 Calotheca , 326-330 Capanemia, 28-33 Capanemia australis , 28-33 Capanemia gehrtii, 28-33 Capanemia hatschhachii , 28-33 Capanemia perpusilla , 28-33 Capanemia therezae , 28-33 Castanopsis, 317-321 Castanopsis glabrifolia, 317-321 Castelnavia , 475-480 Central America, 23-27, 58-65, 94^117, 118-132, 133-148, 413-423, 437-439, 475-480, 483-486, 496-507. see 33-148 Cephae Cephaelis c Cestrum, 244^248 Cestrum oblongifolium, 244-248 Chascolytrum, 326-330 Chascolytrum ambiguum, 326-330 Chascolytrum bidentatum, 326-330 Chascolytrum brasiliense, 326-330 Chascolytrum brizoides, 326-330 Chascolytrum calotheca , 326-330 Chascolytrum itatiaiae, 326-330 Chascolytrum koelerioides , 326-330 Chascolytrum monandrum, 326-330 D Dalbergaria , 413^23 Dalbergieae, 73-77, 331-337 Darcya, 47-48 Darcyanthus, 47^8 Davallia , 380-384 Davallia napoensis , 380-384 ZWZZia w&fcx, 380-384 Davalliaceae, 380-384 Dendrophthora microphylla, 444^462 Dendrophthora subsessilis, 444^462 Dioclea, 226-243 Volume 21, Number 4 Subject Index 523 2011 sect. Macrocarpon, 226-243 ser. Virgatae, 226-243 subg. Pachylobium, 226-243 subg. Platylobium, 226-243 Dioclea apiculata, 226-243 Diodea circinata, 226-243 Dioclea haughtii, 226-243 Dioclea jamesonii, 226-243 Dioclea ovahs , 226-243 Dioclea vallensis, 226-243 Elatostema, 212-215, 281-284 sect. Weddeha ser. Stipulosa, 212-215 Elatostema nasutum. 212-215 Elatostema opposition, 212-215 Enanthecium. 326-330 Ericaceae, 338-342 Euphorbiaceae, 169-173, 192-195, 468-471 Eurasia, 326-330 Fabaceae, 4-19, 226-243, 322-325 Fagaceae, 274-277, 317-321 Fagopyrum, 256-261 Fallopia, 388-391 var. angulata, 388-391 French Guiana, 463-467 Gardenieae. 133-148 Gaidtheria, 338-342 Gaidtheria borneensis, 338-342 Gaidtheria paucinervia, 338-342 Genista dorycnifolia, 4-19 Gouania tihifolia, 481-482 subsp. glandidosa, 481-482 Gramineae, 326-330 Groutiella, 290-293 Groutiella laxotorquata, 290-293 Groutiella pobegumii , 290-293 Groutiella tomentosa, 290-293 Gvmnachne. 326-330 e, 251-255 Hamelieae, 94-117 Heel i, 362-36 Hechtia edulis, 362-367 Hedysarum bifolium , 331-337 Henriquezieae. 118-132 Himalayas, 90-93 Himahnopteris. 90-93 Hqjfmamua. 94-117 Hoffmannia barbillana, 94—117 Hoffmannia boliviano, 94-117 Hoffmannia coriaeea, 94-117 Hoffmannia pearcei, 94—117 Hoffmannia pittien, 94-117 524 Novon Hoffmunmu ieraguens,s. 04-117 Hoya, 508-514 sect. Hoya, 343-346 Hoya fungii, 343-346 Hoya ignorata, 508-514 Hoya jianfenglingensis, 343-346 Hybanthus , 249-250 Hybanthus longistylus, 249-250 Inga pitmanii, 322-325 Iochroma, 491-495 lochrorna fuchsioides , 491-495 Iochrominae. 491-495 Jatropha, 192-195 Jatropha mirandana, 192-195 K Korea, 373-374 Lombardochloa , 326-330 Lomelosia, 402-404 Lomelosia deserticola, 402-404 Lomelosia poecilocarpa, 402-404 Lomelosia transcaspica, 402-404 Loranthaceae, 463-467 sect. Oblonga, 73-77 Machaerium jobimianum, 73-77 Madagascar. 481-482 Magnoliaceae. 375-379 Malaysia, 338-342, 508-514 Manglietia, 375-379 Manglietia crassifolia , 375-379 Manglietia dandyi, 375-379 Manglietia megaphylla, 375-379 Marantaceae. 49-57, 58-65, 66-68 Mascarenes. 481-482 483-486 Chiapas, 393-401 Colima, 483-486 Guerrero, 192-195, 196-200, 244-248 Jalisco, 483-486 Oaxaca, 393-401 Middle America, 393-401. * Musa, 405-412 Musa basjoo var .formosana, 405-412 Musa fonnosana, 405-412 Musa itinerans, 405-412 var. formosana, 405-412 Musareae. 349-350. 405-4K Musci, 424-425 Musella, 349—350 ^ var. rubribraeteata, 349-35 N NautUocalyx dresslen, 413-423 Neotropics, 174-177, 331-337, 357-361, 496-507 Nepal, 90-93 Nicaragua, 94-117, 118-132 Xogopterium. 424-425 Nogopterium beyrichianum , 424—425 Nogoptenum graede, 424-425 North America, 354-356 Volume 21, Number 2011 Subject Index 525 Notopleura, 133-148 Notopleura polyphlebia, 133-148 Notopleura recondita, 133-148 Noveloa, 475-480 0 Odontocarya, 357-361 sect. Somphoxylon, 357-361 Odontocarya arifolia, 357-361 Odontocarya uva-alba, 357-361 Odontochilus, 20-22 Odontochilus tetrapterus, 20-22 Olyreae, 40—43 Olyrineae, 40-43 Oncidiinae, 28-33, 178-181 Orchidaceae, 20-22, 28-33, 178-181 Orthotrichaceae. 200-293 Osmunda, 354-356 Osmundaceae. 354-356 Osmund antrum. 354—356 var. glandulosum, 354-356 Paehycarpus, 426-430 Paehycarpus aeidostelma. 426-430 Paeh ycarpus asperifolius, 426—430 Palicourea, 133-148 Palicourea hammelii. 133-148 Palicourea matamana, 133-148 Palicourea orosiana. 133-148 Palicourea provideneiana, 133-148 toideae, 149-153, 3 ietiferum. 182 Papilionoideae, 331-337 Parodiol yra. 40-43 Paspalum, 368-372 Paspalum Inanstatum. 368-372 Paspalum rostratum , 368-372 Paulluiia. 196-200 Pailllinieae. 196-200 Pentadenia. 413-423 Pentadenia lutea, 413—423 subg. Tildenia. 266-269 Peru, 169-173 Petroeosmea. 385-387 Petrocosmea huanjiangensis. 385-387 Phoradendron, 444-462 Phoradendron alatum, 444—462 Phoradendron camposii. 444-462 Phoradendron concinnum, 444-462 Phoradendron mirandensis , 444-462 Phoradendron nickrentianum, 444—462 Phoradendron palandensis, 444—462 Ph ysoearpus. 373—374 Piperaceae, 266-269 Poa koelerundes. 326-330 Poa tumidula. 326-330 Poaceae, 40-43, 149-153, 326-330, 368 Podostemaceae, 475-480 Poeae, 326-330 Poidium. 326-330 Polygonaceae, 256-261, 388-391 var. angulatum. 388-391 Polypodioideae, 90-93 Pooideae, 326-330 18-132 Posoqueria eorreana. 118-132 Posoqueria panamensis. 118-132 Posoquerieae, 118-132 subsp. boraginoides. 133-148 Psychotria boraginoides, 133-148 Psychotria croatu, 133-148 526 Novon Pterogonium, 424-425 Quercia, 274-277 Quercus delgadoana, 274—277 Quercia salicifolia, 274-277 Quercia sapotifolia, 274—277 Resia, 413-423 Resia bracteata, 413-423 Rhamnaceae. 71-72, 481-482 Rhombolytrum, 326-330 Rhombolytrum berteroanum, 326-330 Rondeletieae. 406-507 Rosaceae, 299-316, 373-374 Rubiaceae, 94-117, 118-132, 133-148, 496-50' Rudgea, 133-148 Rudgea thyrsiflora, 133-148 Salvia, 34-35 subsp. canescens, 34-35 ;, 196-200 Scabiosa, 402-404 Schizocalyx, 496-507 Schizocalyx multiflon Schizocalyx truiicatus, 496-507 Sciaphila , 154-157 Sciaphila arfakiaria, 154-157 Senecio. 347-348 sect. Suffruticosi , 347-348 ser. Chilenses , 347-348 ser. Suffruticosi, 347-348 Simira, 133-148 Simira daricncnsis, 133-148 Solanaceae, 23-27, 47^48, 244-248, 487-490, 491-4 Solarium, 23-27, 487-490 sect. Gcminata, 23-27 subg. Lcptostcnumum sect. Acanthophora, 487—490 Solarium rovirosanurn, 23-27 Solarium savarmarurn, 487-490 subsect. Junceae, 219-225 Solidago juncea, 219-225 South Africa, 426-430 KwaZulu-Natal, 426-430 South America, 94-117, 118-132, 219-225, 326-330, , 348, 413-423, 483-486, 487-490, 496-507. see Southeast Asia, 508-514 sect. Glomeratac, 299-316 / adiantoides, 299-316 / compsophylla, 299-316 / lasiocarpa, 299-316 villosa, 299-316 / media, 299-316 ongolica 99-316 •a sericea, 299-316 tarpheti ■ustifolia. Strychiios, 472—474 Strychiios guianensis, 472-474 P , 357-361 Volume 21, Number 2011 Subject Index 527 Tetrorchidium trichotocarpum, 468-471 Thailand, 149-153, 508-514 Vietnam, 375-379, 508-514 Turkey, 34-35, 182 Anatolia, 34-35 Zingiberaeeae. 270-273 Zornia, 331-337 Zornia diphylla, 331—337" f. cUiata, 331-337 var. bernardinensis, 331-337 Venezuela, 40-43, 178-181, 183-186, 413-423, 444-462, 475-480 Amazonas, 178-181, 183-186, 475-480 Volume 21, Number 1, pp. 1-160 of NOVON was published on 7 April 2011. Volume 21, Number 2, pp. 161-294 of NOVON was published on 27 June 2011. Volume 21, Number 3, pp. 295-392 of NOVON was published on 9 September 2011. Volume 21, Number 4, pp. 393-528 of NOVON was published on 29 December 2011. www. mbgpr ess . info CONTENTS I